Sample records for brachiopod obolus apollinis

  1. Late Ordovician brachiopods from eastern North Greenland

    DEFF Research Database (Denmark)

    Mac Ørum Rasmussen, Christian


    Late Ordovician rhynchonelliformean brachiopods, typical of the North American Red River fauna, are found sporadically in the BOrglum River Formation of the Centrum SO area, Kronprins Christian Land, eastern North Greenland. The geographical distribution of this characteristic brachiopod fauna......) to younger strata exclusively yielding specimens of H.gigas. As H.gigas occurs in the upper part of the Cape Calhoun Formation in Washington Land, it indicates that the upper boundary of the Cape Calhoun Formation is considerably younger than previous estimates, reaching into the uppermost Katian (middle...... (Richmondian), it possesses a strong provincial signal during the later Ordovician. The new occurrences indicate that this fauna extended to the north-eastern margin of the Laurentian Craton. It lived in close association with cosmopolitan faunal elements that may have been the earliest sign of the succeeding...

  2. Non-terrestrial food source for Fiordland brachiopods

    International Nuclear Information System (INIS)

    Lyon, G.L.; Richardson, Joyce


    Carbon-13 analyses were determined for brachiopods and particulate organic matter from Fiordland waters. Brachiopod delta 13 $ 0 C are about -18 per mille which is significantly enriched in 13 C relative to the particulate matter (about -23 per mille) and different from local terrestrial matter (about -28 per mille). There is no carbon-13 evidence for non-marine food in the diet of brachiopods


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    Full Text Available Late Wordian (Guadalupian brachiopods from Member 2 of the Panjshah Formation in the Karakorum (N Pakistan are described. The brachiopod assemblage, dated by the associated fusulinids and conodonts, consists of 29 genera (3 of which are questionable and 1 unidentifiable of the orders Productida, Orthida, Rhynchonellida, Athyridida, Spiriferida and Terebratulida. Hunzininae, a new subfamily of the Spiriferellidae is proposed; it includes Darbandia n. gen., with type species D. vagabunda n. sp. and Elivina chapursani n. sp. A third new species is assigned to the genus Anchorhynchia of the family Wellerellidae: A. cimmerica n. sp.A quantitative biostratigraphic analysis demonstrates two major faunal changes in the Elivina chapursani- Chapursania tatianae Assemblage Zone of the upper part of Member 2, which are not strictly linked to lithological changes. This biozone is correlated with the brachiopod faunas of the Gnishik Formation of Armenia and those of the basal Takhtabulak Formation of SE Pamir.The faunal elements of the Elivina chapursani- Chapursania tatianae Assemblage Zone are an admixture of wide-ranging, Tethyan (particularly abundant, Gondwanan and endemic (Cimmerian genera, representing a transitional fauna and a biostratigraphic tool for intercontinental correlation, which are particularly problematic in this time interval. The Panjshah transitional fauna demonstrates the persistence of the Transhimalayan Province of the Cimmerian Region into the late Guadalupian, which originated at the end of the Cisuralian and occupied Armenia, Central Afghanistan, Karakorum and SE Pamir. It provides also some insights into the biodiversity pattern before the mass extinction at the end of the Guadalupian, and suggests that this event was as rapid as the end- Permian mass extinction, at least in Central Asia. 

  4. Phylogenetic paleobiogeography of Late Ordovician Laurentian brachiopods

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    Jennifer E. Bauer


    Full Text Available Phylogenetic biogeographic analysis of four brachiopod genera was used to uncover large-scale geologic drivers of Late Ordovician biogeographic differentiation in Laurentia. Previously generated phylogenetic hypotheses were converted into area cladograms, ancestral geographic ranges were optimized and speciation events characterized as via dispersal or vicariance, when possible. Area relationships were reconstructed using Lieberman-modified Brooks Parsimony Analysis. The resulting area cladograms indicate tectonic and oceanographic changes were the primary geologic drivers of biogeographic patterns within the focal taxa. The Taconic tectophase contributed to the separation of the Appalachian and Central basins as well as the two midcontinent basins, whereas sea level rise following the Boda Event promoted interbasinal dispersal. Three migration pathways into the Cincinnati Basin were recognized, which supports the multiple pathway hypothesis for the Richmondian Invasion.

  5. The oldest brachiopods from the lower cambrian of South Australia

    DEFF Research Database (Denmark)

    Topper, Timothy Paul; Holmer, Lars E.; Skovsted, Christian B.


    The morphology and organophosphatic shell structure of the paterinate brachiopod Askepasma is documented using new and previously collected specimens from the lower Cambrian of South Australia. Lack of adequately preserved material has seen the majority of paterinate specimens previously reported...... from South Australia referred to the genus Askepasma and treated under open nomenclature. Large collections of paterinates from the lower Cambrian Wilkawillina, Ajax, and Wirrapowie limestones in the Arrowie Basin, South Australia have prompted redescription of the type species Askepasma toddense...... and the erection of a new species, Askepasma saproconcha sp. nov. Askepasma saproconcha sp. nov. currently represents the oldest known brachiopod from the lower Cambrian successions in South Australia with a FAD in pre-trilobitic (Terreneuvian, Cambrian Stage 2, lower Atdabanian) strata in the basal part...

  6. Early Ordovician (Tremadocian brachiopods from the Eastern Alborz Mountains, Iran

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    Leonid E. Popov


    Full Text Available Six linguliform and two rhynchonelliform brachiopods, including three new species Eurytreta ahmadii, Wahwahlingula kharbashi and Nanorthis bastamensis are described from Tremadocian strata (Paltodus deltifer deltifer conodont Biozone in the Deh-Molla area southwest of Shahrud, Northern Iran. The fauna is dominated by micromorphic lingulides and acrotretides and shows distinct similarity to the contemporaneous micromorphic brachiopod association from Tremadocian chalcedonites of the Holy Cross Mountains, Poland. New data on the early ontogeny of the enigmatic lingulide Diencobolus show a very distinct pattern, including the presence of a metamorphic protegulum ornamented with flat-based pits and a single pair of larval setal bundles, which links this taxon to Paterula and suggests close phylogenetic relationships of both taxa to the Discinoidea.

  7. Middle Ordovician brachiopods from the Stairway Sandstone, Amadeus Basin, central Australia

    DEFF Research Database (Denmark)

    Jakobsen, Kristian Grube; Brock, Glenn A.; Nielsen, Arne Thorshøj


    -water palaeoenvironment along the margin of northeastern Gondwana. The brachiopods from the Stairway Sandstone are of low diversity and represent ca 9% of the entire shelly fauna. Five brachiopod taxa are described from the Stairway Sandstone; all are endemic to the Amadeus Basin at species level. Two new species...

  8. Biodiversity, biogeography and phylogeography of Ordovician rhynchonelliform brachiopods

    DEFF Research Database (Denmark)

    Harper, David A. T.; Mac Ørum Rasmussen, Christian; Liljeroth, Maria


    -levels were high. Pivotal to the entire diversification is the role of gamma (inter-provincial) diversity and by implication the spread of the continents and frequency of island arcs and microcontinents. The phylogeographical analysis demonstrates that this new palaeogeographical configuration...... with local biodiversity epicentres, notably on the South China Palaeoplate; low-latitude porambonitoid-dominated faunas with early plectambonitoid and clitambonitoid representatives, as well as high-latitude assemblages mostly dominated by orthoids, can be recognized, but many taxa are rooted in Late......The phylogeographical evolution and the consequent changing distribution and diversity of rhynchonelliform brachiopods through the Ordovician are linked to the dynamic palaeogeography of the period. The Early Ordovician (Tremadocian and Floian) is characterized by globally low-diversity faunas...

  9. Brachiopod biogeochemistry and isotope stratigraphy from the Rhaetian Eiberg section in Austria

    DEFF Research Database (Denmark)

    Korte, Christoph; Thibault, Nicolas Rudolph; Ullmann, Clemens Vinzenz


    The end-Triassic is characterized by one of the most severe biotic crises of the entire Phanerozoic, with strong carbon cycle perturbations potentially predating the biotic event. In order to improve and test the chemostratigraphic framework for the Rhaetian Stage, which culminated in the end......-Triassic extinction, a total of 675 and 108 carbonate δ13C and δ18O values have been measured from bulk rock carbonate and articulate brachiopod samples from the Rhaetian Eiberg quarry succession (Northern Calcareous Alps, Austria) respectively. Both brachiopod and bulk rock oxygen isotope trends are compatible...... the calcite composing the bulk rock micrite. Significant metabolic disequilibrium effects on brachiopod carbon isotope ratios are suggested by a clear negative correlation between δ13C values and Sr/Ca ratios of the samples. Consequently, chemostratigraphy based on brachiopod Δ13C data, especially with regard...

  10. Ecological interactions on macroevolutionary time scales: clams and brachiopods are more than ships that pass in the night. (United States)

    Liow, Lee Hsiang; Reitan, Trond; Harnik, Paul G


    Competition among organisms has ecological and evolutionary consequences. However, whether the consequences of competition are manifested and measureable on macroevolutionary time scales is equivocal. Marine bivalves and brachiopods have overlapping niches such that competition for food and space may occur. Moreover, there is a long-standing debate over whether bivalves outcompeted brachiopods evolutionarily, because brachiopod diversity declined through time while bivalve diversity increased. To answer this question, we estimate the origination and extinction dynamics of fossil marine bivalve and brachiopod genera from the Ordovician through to the Recent while simultaneously accounting for incomplete sampling. Then, using stochastic differential equations, we assess statistical relationships among diversification and sampling dynamics of brachiopods and bivalves and five paleoenvironmental proxies. None of these potential environmental drivers had any detectable influence on brachiopod or bivalve diversification. In contrast, elevated bivalve extinction rates causally increased brachiopod origination rates, suggesting that bivalves have suppressed brachiopod evolution. © 2015 The Authors. Ecology Letters published by CNRS and John Wiley & Sons Ltd.

  11. Population structure, growth and production of a recent brachiopod from the Chilean fjord region

    KAUST Repository

    Baumgarten, Sebastian


    Magellania venosa, the largest recent brachiopod, occurs in clusters and banks in population densities of up to 416 ind m-2 in Comau Fjord, Northern Chilean fjord region. Below 15 m, it co-occurs with the mytilid Aulacomya atra and it dominates the benthic community below 20 m. To determine the question of why M. venosa is a successful competitor, the in situ growth rate of the brachiopod was studied and its overall growth performance compared with that of other brachiopods and mussels. The growth in length was measured between February 2011 and March 2012 after mechanical tagging and calcein staining. Settlement and juvenile growth were determined from recruitment tiles installed in 2009 and from subsequent photocensus. Growth of M. venosa is best described by the general von Bertalanffy growth function, with a maximum shell length (L∞) of 71.53 mm and a Brody growth constant (K) of 0.336 year-1. The overall growth performance (OGP index = 5.1) is the highest recorded for a rynchonelliform brachiopod and in the range of that for Mytilus chilensis (4.8-5.27), but lower than that of A. atra (5.74). The maximal individual production (PInd) is 0.29 g AFDM ind-1 year-1 at 42 mm shell length and annual production ranges from 1.28 to 89.25 g AFDM year-1 m-2 (1-57% of that of A. atra in the respective fjords). The high shell growth rate of M. venosa, together with its high overall growth performance may explain the locally high population density of this brachiopod in Comau Fjord. However, the production per biomass of the population (P/B--ratio) is low (0.535) and M. venosa may play only a minor role in the food chain. Settling dynamics indicates that M. venosa is a pioneer species with low juvenile mortality. The coexistence of the brachiopod and bivalve suggests that brachiopod survival is affected by neither the presence of potential brachiopod predators nor that of space competitors (i.e. mytilids).

  12. Population structure, growth and production of a recent brachiopod from the Chilean fjord region

    KAUST Repository

    Baumgarten, Sebastian; Laudien, Jü rgen; Jantzen, Carin; Hä ussermann, Verena; Fö rsterra, Gü nter


    -57% of that of A. atra in the respective fjords). The high shell growth rate of M. venosa, together with its high overall growth performance may explain the locally high population density of this brachiopod in Comau Fjord. However, the production per biomass

  13. Brachiopod associations from the Middle Ordovician of the Oslo Region, Norway

    DEFF Research Database (Denmark)

    Candela, Yves; Hansen, Thomas


    The marine upper Darriwilian (Middle Ordovician) Elnes Formation of southern Norway contains very rich and diverse invertebrate faunas. Stratigraphically detailed recent collections of these well-preserved faunas have permitted a more thorough description of the various faunal groups and their pr......The marine upper Darriwilian (Middle Ordovician) Elnes Formation of southern Norway contains very rich and diverse invertebrate faunas. Stratigraphically detailed recent collections of these well-preserved faunas have permitted a more thorough description of the various faunal groups...... by linguliformeans although some sections of the formation are dominated by the plectambonitoids Alwynella ildjernensis Spjeldnaes or Cathrynia aequistriata (Hadding) or by orthid brachiopods. Statistical analysis of the brachiopod fauna reveals that the general depositional trend of the Elnes Formation...... in muddy to marly deposits formed around maximal storm wave base and an orthid-dominated association representing an environment characterised by a fairly coarse or hard bottom substrate formed well above storm wave base but below fair weather wave base....

  14. Late Ordovician palaeogeography and the positions of the Kazakh terranes through analysis of their brachiopod faunas (United States)

    Popov, Leonid E.; Cocks, Robin M.


    Detailed biogeographical and biofacies analyses of the Late Ordovician brachiopod faunas with 160 genera, grouped into 94 faunas from individual lithotectonic units within the Kazakh Orogen strongly support an archipelago model for that time in that area. The Kazakh island arcs and microcontinents within several separate clusters were located in the tropics on both sides of the Equator. Key units, from which the Late Ordovician faunas are now well known, include the Boshchekul, Chingiz-Tarbagatai, and Chu-Ili terranes. The development of brachiopod biogeography within the nearly ten million year time span of the Late Ordovician from about 458 to 443 Ma (Sandbian, Katian, and Hirnantian), is supported by much new data, including our revised identifications from the Kazakh Orogen and elsewhere. The Kazakh archipelago was west of the Australasian segment of the Gondwana Supercontinent, and relatively near the Tarim, South China and North China continents, apart from the Atashu-Zhamshi Microcontinent, which probably occupied a relatively isolated position on the south-western margin of the archipelago. Distinct faunal signatures indicate that the Kazakh terranes were far away from Baltica and Siberia throughout the Ordovician. Although some earlier terranes had joined each other before the Middle Ordovician, the amalgamation of Kazakh terranes into the single continent of Kazakhstania by the end of the Ordovician is very unlikely. The Late Ordovician brachiopods from the other continents are also compared with the Kazakh faunas and global provincialisation statistically determined.

  15. Late Permian marine ecosystem collapse began in deeper waters: evidence from brachiopod diversity and body size changes. (United States)

    He, W-H; Shi, G R; Twitchett, R J; Zhang, Y; Zhang, K-X; Song, H-J; Yue, M-L; Wu, S-B; Wu, H-T; Yang, T-L; Xiao, Y-F


    Analysis of Permian-Triassic brachiopod diversity and body size changes from different water depths spanning the continental shelf to basinal facies in South China provides insights into the process of environmental deterioration. Comparison of the temporal changes of brachiopod diversity between deepwater and shallow-water facies demonstrates that deepwater brachiopods disappeared earlier than shallow-water brachiopods. This indicates that high environmental stress commenced first in deepwater settings and later extended to shallow waters. This environmental stress is attributed to major volcanic eruptions, which first led to formation of a stratified ocean and a chemocline in the outer shelf and deeper water environments, causing the disappearance of deep marine benthos including brachiopods. The chemocline then rapidly migrated upward and extended to shallow waters, causing widespread mass extinction of shallow marine benthos. We predict that the spatial and temporal patterns of earlier onset of disappearance/extinction and ecological crisis in deeper water ecosystems will be recorded during other episodes of rapid global warming. © 2014 John Wiley & Sons Ltd.

  16. Neuromuscular development in Novocrania anomala: evidence for the presence of serotonin and a spiralian-like apical organ in lecithotrophic brachiopod larvae

    DEFF Research Database (Denmark)

    Altenburger, Andreas; Wanninger, Andreas Wilhelm Georg


    SUMMARY The phylogenetic position of Brachiopoda remains unsettled, and only few recent data on brachiopod organogenesis are currently available. In order to contribute data to questions concerning brachiopod ontogeny and evolution we investigated nervous and muscle system development in the cran......SUMMARY The phylogenetic position of Brachiopoda remains unsettled, and only few recent data on brachiopod organogenesis are currently available. In order to contribute data to questions concerning brachiopod ontogeny and evolution we investigated nervous and muscle system development...... stages. In the juveniles, the nervous system stained by a-tubulin is characterized by two ventral neurite bundles with three commissures. Our data are the first direct proof for the presence of an immunoreactive neurotransmitter in lecithotrophic brachiopod larvae and demonstrate the existence of flask...

  17. Resolving early Mid-Ordovician (Kundan) bioevents in the East Baltic based on brachiopods

    DEFF Research Database (Denmark)

    Rasmussen, Christian Mac Ørum; Harper, David Alexander Taylor


    Bed-by-bed sampling of brachiopods from selected Kundan Stage localities in the East Baltic has permitted the construction of a-diversity curves, unprecedented in detail from this particular stage and region. These a-diversity curves can be correlated between the investigated sections, along...... an east to west transect of some 200 km and indicate bioevents associated with intervals of increased palaeo-water depth. The highest peaks in a-diversity occur when biofacies analysis and sedimentological evidence indicate drowning events. The events involve mainly taxa endemic to the Baltic province...

  18. Element/Ca, C and O isotope ratios in modern brachiopods

    DEFF Research Database (Denmark)

    Ullmann, C. V.; Frei, Robert; Korte, Christoph


    Fossil brachiopods are of major importance for the reconstruction of palaeoenvironmental conditions, particularly of the Paleozoic. In order to better understand signals of ancient shell materials, modern analogue studies have to be conducted. Here we present C and O isotope data in conjunction...... with ambient seawater. Some species – especially in the suborder Terebratellidina – show partly distinct disequilibrium signals, suggesting some degree of phylogenetic control on the expression of vital effects. Mn/Ca and Fe/Ca ratios measured in the modern species form a baseline to assess fossil preservation...

  19. 87Sr/86Sr ratios in permo-carboniferous sea water from the analyses of well-preserved brachiopod shells

    International Nuclear Information System (INIS)

    Popp, B.N.; Anderson, T.F.


    Sr isotopic analyses of well-preserved portions of Permo-Carboniferous brachiopods distributed globally confirm the general shape of the Sr isotope age curve established by previous workers for this time interval. There is little variation between the SR isotopic composition of unaltered portions of brachiopods and that of portions of the same shell interpreted to be diagenetically altered (based on cathodoluminescence, elemental, and stable isotopic data). However, the Sr isotopic composition in diagenetically altered micritic matrix adjacent to the shell is more radiogenic. The Sr isotopic composition in the unaltered portions of calcitic megafossils has potential as a stratigraphic tool. (author)

  20. Synchrotron radiation X-ray tomographic microscopy (SRXTM of brachiopod shell interiors for taxonomy: Preliminary report

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    Motchurova-Dekova Neda


    Full Text Available Synchrotron radiation X-ray tomographic microscopy (SRXTM is a non-destructive technique for the investigation and visualization of the internal features of solid opaque objects, which allows reconstruction of a complete three-dimensional image of internal structures by recording of the differences in the effects on the passage of waves of energy reacting with those structures. Contrary to X-rays, produced in a conventional X-ray tube, the intense synchrotron light beams are sharply focused like a laser beam. We report encouraging results from the use of SRXTM for purely taxonomic purposes in brachiopods: an attempt to find a non-destructive and more efficient alternative to serial sectioning and several other methods of dissection together with the non-destructive method of X-ray computerised micro-tomography. Two brachiopod samples were investigated using SRXTM. In “Rhynchonella” flustracea it was possible to visualise the 3D shape of the crura and dental plates. In Terebratulina imbricata it was possible to reveal the form of the brachidium. It is encouraging that we have obtained such promising results using SRXTM with our very first two fortuitous samples, which had respectively fine-grained limestone and marl as infilling sediment, in contrast to the discouraging results communicated to us by some colleagues who have tested specimens with such infillings using X-ray micro-tomography. In future the holotypes, rare museum specimens or delicate Recent material may be preferentially subjected to this mode of analysis.

  1. Brachiopod faunas after the end Ordovician mass extinction from South China: Testing ecological change through a major taxonomic crisis (United States)

    Huang, Bing; Harper, David A. T.; Rong, Jiayu; Zhan, Renbin


    Classification of extinction events and their severity is generally based on taxonomic counts. The ecological impacts of such events have been categorized and prioritized but rarely tested with empirical data. The ecology of the end Ordovician extinction and subsequent biotic recovery is tracked through abundant and diverse brachiopod faunas in South China. The spatial and temporal ranges of some 6500 identified specimens, from 10 collections derived from six localities were investigated by network and cluster analyses, nonmetric multidimensional scaling and a species abundance model. Depth zonations and structure of brachiopod assemblages along an onshore-offshore gradient in the late Katian were similar to those in the latest Ordovician-earliest Silurian (post-extinction fauna). Within this ecological framework, deeper-water faunas are partly replaced by new taxa; siliciclastic substrates continued to be dominated by the more 'Ordovician' orthides and strophomenides, shallow-water carbonate environments hosted atrypides, athyridides and pentamerides, with the more typical Ordovician brachiopod fauna continuing to dominate until the late Rhuddanian. The end Ordovician extinctions tested the resilience of the brachiopod fauna without damage to its overall ecological structure; that commenced later at the end of the Rhuddanian.

  2. A bradoriid and brachiopod dominated shelly fauna from the furongian (Cambrian) of Västergötland, Sweden

    DEFF Research Database (Denmark)

    Topper, Timothy Paul; Skovsted, Christian B.; Harper, David Alexander Taylor


    A small assemblage of shelly fossils, dominated by the brachiopod Treptotreta jucunda and the bradoriid arthropod Mongolitubulus aspermachaera new species is described from a Furongian limestone of Västergötland, south-central Sweden. Mongolitubulus aspermachaera is represented in the assemblage ...... include typical Baltic Furongian representatives, such as the trilobite Parabolina, the agnostoid Agnostus and the phosphatocopids Hesslandona and Vestrogothia....

  3. Brachiopods and rugose corals in an upper Serpukhovian (Mississippian) biostrome: preliminary results from the Djebel Arhlal (Bechar Basin, Algeria)

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    Atif, K. F. T.; Aretz, M.; Legrand-Biain, M.; Bouzidi, A.; Aimouche, M.


    The Djebel Arhlal is the southernmost outcrop of upper Serpukhovian strata in the Bechar Basin. Here the Djenien member of the Djenien Formation is three-folded and it contains a 10 m-thick coral biostrome in its middle unit formed by Siphonodendron, Diphyphyllum and Lithostrotion. Distances between in situ colonies are in the order of several decimetres and the space between them is filled with bioclastic limestone containing coral rubble. Few brachiopods and solitary corals are found as dwellers within the biostrome, but these groups are much more common in the strata below and above the coral biostrome. This is especially true for the brachiopods, which reach diversities of more than a dozen species in specific horizons. They mainly belong to the orders Productida, Spiriferida and Athyridida. Agitated open-marine platform interior or platform margin settings are the general facies encountered in the Djenien member at the Djebel Arhlal. The field data, including brachiopod coquinas and many fragmented brachiopods and corals, are confirmed by microfacies analysis. However, the coral biostrome records a quieter setting at the interface of middle and outer ramp settings, as seen in the co-existence of in situ coral growth, input of reworked material, deposition of carbonate mud, and sparitic textures. This autoparabiostrome at Djebel Arhlal is compared to other rather thick and of exceptional horizontal extension upper Serpukhovian biostromes (few kilometres). (Author)

  4. Microstructural growth increments in the brachiopods Liothyrella uva and L. neozelanica: preliminary study of growth analysis and proxy calibration (United States)

    Romanin, Marco; Aleksandra Bitner, Maria; Brand, Uwe


    Brachiopods secrete low-Mg calcite shells in near equilibrium with the surrounding sea water, with respect to their secondary and tertiary layers. For this reason, in recent years they have been intensively studied as archives for oceanographic and environmental proxies. The primary layer has been shown not to be deposited in equilibrium with the ambient sea water, leading to a novel cleaning protocol proposed by Zaki et al (2015). In the spite of improving on existing proxies, the shell microstructure and growth has to be taken in to account in their applications. The secretion of the primary layer is known to be external of the shell, but in SEM investigations of Liothyrella uva and L. neozelanica we discovered that the primary layer has its origin within the fibres of the secondary layer. Furthermore, the primary layer calcite is not a continuum but instead it consists of a 'new' band for each major growth increment. There is overlap between the preceding and subsequent 'band' (or shingles) of the primary layer, which may extend into the secondary/tertiary layer. This finding may lead to more comprehensive knowledge of shell microstructure processes in L. uva and L. neozelanica that may be applied and extended to other modern and fossil brachiopods, including age dating of brachiopods. This discovery may make brachiopod archives more reliable and consistent proxies when applied to and interpreting their geological record.

  5. Mollusc and brachiopod skeletal hardparts: Problematic archives of past seawater properties (Invited) (United States)

    Immenhauser, A.; Schone, B. R.; Hoffmann, R.; Niedermayr, A.


    Biomineralized exo- or endoskeletons of fossil marine invertebrates are widespread and diverse components of the Phanerozoic rock record of Earth's past and present oceans. Exoskeletons serve as protection against environmental pressure or predators, whilst endoskeletons may act as support or serves as an attachment for muscles and ligaments and hence as a mechanism for transmitting muscular forces. Biogenic hardparts represent sophisticated products resulting from the hierarchical interaction of inorganic minerals (95%) and macromolecular organic matrix, forming commonly less than 5%. The significance of biogenic carbonate archives lies in the time-resolved growth-increments and their respective multi-proxy geochemical signatures that have been widely used to assess past oceanic seawater properties. We here compile and review published work dealing with crystallization pathways of skeletal hardparts secreted by molluscs (i.e., bivalves and cephalopods) as well as brachiopods as widely used archives of ancient neritic epeiric settings. Bivalves and cephalopods (e.g., extinct ammonoids and belemnites and extant Sepia, Nautilus and Spirula) all form accretionary calcitic, aragonitic or vateritic skeletal hardparts. Despite the fact that molluscs and brachiopods form part of very different branches of the animal phylogenetic tree, their biomineralization strategies are surprisingly similar. Our main focus lies in a critical assessment of the complex pathways of ions and aquo-complexes from their source (seawater) to the final product (biomineral). We do this as an attempt to critically test the commonly held hypothesis that many fossil shell hardparts precipitated, under favourable conditions and pending subsequent diagenetic alteration, in equilibrium with seawater. Two main observations stand out: (1) the present knowledge on pathways and mechanisms (e.g., ion channel trans-membrane or endocytosis and vesicle transport, precursor mineralogies etc.) is surprisingly

  6. Decadal climate variation recorded in modern global carbonate archives (brachiopods, molluscs) (United States)

    Romanin, Marco; Zaki, Amir H.; Davis, Alyssa; Shaver, Kristen; Wang, Lisha; Aleksandra Bitner, Maria; Capraro, Luca; Preto, Nereo; Brand, Uwe


    The progress of the Earth's warming trend has rapidly accelerated in the last few decades due to the increase in emission of anthropogenic greenhouse gases. The exchange of heat between the atmosphere and seawater has consequently elevated the rate of temperature buildup in the low and high latitude ocean. Records of the variation in seawater temperature in response to local and global changes in climate are preserved within the carbonate structures of marine biogenic archives. Investigating the isotopic composition of the archives' growth increments documents the magnitude of sea surface temperature (SST) change. A long-term (1956-2012) record of temperature change in sub-tropical seawater was acquired from the giant clam Tridacna maxima collected from the Red Sea in conjunction with published results of the oyster Hyotissa hyotis (Titschack et al., 2010). Variation in polar-subpolar SST was obtained from the brachiopod Magellania venosa recovered from the coastal area of southern Chile, and from the proxy record of Hemithiris psittacea of Hudson Bay (Brand et al., 2014). The former reveals a long-term (1961-2012) time-series of Antarctic-induced oceanographic change in the southern hemisphere, while the latter represents a trend of Hudson Bay seawater SST in the northern hemisphere. Evaluation of the isotopic compositions confirms the equilibrium incorporation of oxygen isotopes with respect to ambient seawater in brachiopods and some bivalves. A general trend of decreasing δ18O values in the Red Sea molluscs is observed, indicating an increase in tropical seawater temperature of about 0.79°C since 1988. The δ18O values of the polar-subpolar brachiopods display similar depletion slopes but of larger magnitudes than that of the Red Sea archives. This signifies a rise in seawater temperature of about 1.47°C in Hudson Bay since 1991, and about 2.08°C in southern Chile since 1988. The 2013 IPCC report suggests an increase in SST of +0.094°C per decade (average

  7. The eoorthid brachiopod Apheoorthina in the Lower Ordovician of NW Argentina and the dispersal pathways along western Gondwana

    Directory of Open Access Journals (Sweden)

    Diego F. Muñoz


    Full Text Available The eoorthid brachiopod Apheoorthina is reported for the first time from the Lower Ordovician of NW Argentina. It is represented by a species similar to A. ferrigena from the Tremadocian of the Prague Basin, increasing the faunal affinities between the Central Andean Basin and the South European microcontinents, in particular the Bohemian region (Perunica. Nine out of the fourteen brachiopod genera reported from the Tremadocian of the Central Andean Basin (~64% are shared with the Mediterranean region, four of which (~28% have been recorded in the Prague Basin, and two (Kvania and Apheoorthina are restricted to the Central Andes and Perunica. Dispersal pathways around Gondwana are analyzed in the light of major factors affecting large-scale distribution of brachiopods (environment, larval capacity for dispersal, oceanic currents. The presence in Apheoorthina aff. ferrigena of a well-preserved larval protegulum measuring 420 μm in width and 210 μm in length strongly suggests that this species had planktotrophic larvae capable of long-distance dispersal. According to recent ocean-atmosphere general circulation models for the Ordovician Period, the Central Andean margin was dominated by the cold-water Antarctica Current. Despite the complex non-zonal pattern produced by current deflections around the peri-Gondwanan microcontinents, the general westward circulation sense favoured larval dispersal from the Andean region to North Africa, Avalonia, the Armorican Terrane Assemblage, and Perunica. On the other hand, the eastwards flowing Gondwana Current connected the North Gondwana waters with the South American epicontinental seas, which could explain the reversed migration of some brachiopods.

  8. Platystrophia (Orthida and new related Ordovician and Early Silurian brachiopod genera

    Directory of Open Access Journals (Sweden)

    Harper, David A. T.


    Full Text Available More than 150 Ordovician and Early Silurian brachiopod species have been assigned to the genus Platystrophia King, 1850 mainly on the basis of their Spirifer-like shell exteriors. King’s concept of the genus was based on Platystrophia biforata King, which is not conspecific with Terebratulites biforatus Schlotheim, traditionally regarded as the type species of Platystrophia. Porambonites costatus Pander, 1830 is formally proposed as the type species of the genus to replace P. biforata; the latter is considered to be a nomen dubium. In our revised diagnosis, Platystrophia is restricted to a group of Arenig to upper Caradoc species from Baltica and Avalonia, whereas the Ashgill and lower Silurian taxa of these regions, hitherto assigned to Platystrophia, are placed in the new genus Neoplatystrophia. Platystrophia ponderosa Foerste, 1909 from the Upper Ordovician of North America is proposed as the type species of a new genus Vinlandostrophia. Two new species, Platystrophia baltica and Platystrophia pogrebovi from the Llanvirn–Caradoc of the East Baltic are also described.

  9. Assessing the completeness of the fossil record using brachiopod Lazarus taxa (United States)

    Gearty, W.; Payne, J.


    Lazarus taxa, organisms that disappear from the fossil record only to reappear later, provide a unique opportunity to assess the completeness of the fossil record. In this study, we apply logistic regression to quantify the associations of body size, geographic extent, and species diversity with the probability of being a Lazarus genus using the Phanerozoic fossil record of brachiopods. We find that both the geographic range and species diversity of a genus are inversely associated with the probability of being a Lazarus taxon in the preceding or succeeding stage. In contrast, body size exhibits little association with the probability of becoming a Lazarus taxon. A model including species diversity and geographic extent as predictors performs best among all combinations examined, whereas a model including only shell size as a predictor performs the worst - even worse than a model that assumes Lazarus taxa are randomly drawn from all available genera. These findings suggest that geographic range and species richness data can be used to improve estimates of extensions on the observed fossil ranges of genera and, thereby, better correct for sampling effects in estimates of taxonomic diversity change through the Phanerozoic.

  10. Taxonomic diversity dynamics of early cretaceous brachiopods and gastropods in the Azerbaijanian domains of the Lesser Caucasus (Neo-Tethys Ocean

    Directory of Open Access Journals (Sweden)

    Ruban Dmitry A.


    Full Text Available Palaeontological data available from the Azerbaijanian domains (Somkhit-Agdam, Sevan-Karabakh, and Miskhan-Kafan tectonic zones of the Lesser Caucasus permit reconstruction of the regional taxonomic diversity dynamics of two groups of Early Cretaceous marine benthic invertebrates. Stratigraphical ranges of 31 species and 14 genera of brachiopods and 40 species and 31 genera of gastropods are considered. The total number of species and genera of brachiopods was low in the Berriasian-Valanginian and then rose to peak in the Barremian. Then, the diversity declined in the Aptian, and brachiopods are not known regionally from the Albian. Gastropods appeared in the Hauterivian and experienced a strong radiation in the Barremian. The diversity of species and genera declined in the Aptian (with a minor radiation in the Middle Aptian, and no gastropods are reported from the Albian. Globally, the number of brachiopod genera remained stable through the Early Cretaceous, and the number of gastropod genera increased stepwise with the maximum in the Albian. The regional and global patterns of the diversity dynamics differed for the both groups of marine benthic invertebrates. The Barremian maximum of the taxonomic diversity coincided with the regional flourishing of reefal ecosystems. The taxonomic diversity dynamics of brachiopods in the Azerbaijanian domains of the Lesser Caucasus is very similar to those of the Northern Caucasus, which is an evidence of proximity of these regions during the Early Cretaceous.

  11. Do the Available Data Permit Clarifcation of the Possible Dependence of Palaeozoic Brachiopod Generic Diversity Dynamics on Global Sea-Level Changes? A Viewpoint

    Directory of Open Access Journals (Sweden)

    Ruban Dmitry A.


    Full Text Available At a glance, progress in palaeontology and eustatic reconstructions in the past decade permits to prove or to disprove the possible dependence of Palaeozoic brachiopod generic diversity dynamics on global sea-level changes. However, the available diversity curve is of much lower resolution than the eustatic curve. This problem can be resolved by decreasing the resolution of the latter. The other restriction linked to the chronostratigraphical incompatibility of the available data allows to focus on the Middle Palaeozoic only. A series of mass extinctions and other biotic crises in the Silurian-Devonian does not allow to interpret correctly the results of direct comparison of the brachiopod generic diversity dynamics with global sea-level changes. With the available data, it is only possible to hypothesize that the eustatic control was not playing a major part in diversity dynamics of Middle Palaeozoic brachiopods. The resolution of the stratigraphic ranges of Palaeozoic brachiopods should be increased signifcantly, and these ranges should be plotted against the most up-to-date geologic time scale. Until this task will be achieved, it is impossible to judge about the existence of any dependence (either full or partial of the Palaeozoic brachiopod diversity dynamics on global sea-level changes.

  12. Brachiopods and stratigraphy of the Upper Devonian (Frasnian) succession of the Radlin Syncline (Holy Cross Mountains, Poland) (United States)

    Baliński, Andrzej; Racki, Grzegorz; Halamski, Adam T.


    The lower part of the Frasnian succession in the Radlin Syncline (Kielce-Łagów Synclinorium, southern region of the Holy Cross Mountains), in the two studied successions: Józefka at Górno and (for the first time) Radlin, consists of the rhythmic marly Szydłówek Beds, the fossil-rich limestones of the Wietrznia Beds (locally) and the atypically developed, calcareous Kostomłoty Beds. The carbon isotope chemostratigraphic pattern overall corresponds well to the global Early-Middle Frasnian biogeochemical perturbation, even if the major punctata positive excursion is only fragmentarily recorded in the Kostomłoty intrashelf basin. Two brachiopod assemblages are abundantly represented in both sections: the Phlogoiderhynchus polonicus Assemblage, typical of the Szydłówek Beds, and the Biernatella lentiformis Assemblage, limited to the middle part of the Wietrznia Beds. Both are highly dominated by the index species. Twenty nine lower Frasnian brachiopod species (Craniida - 1 species, Strophomenida - 1, Productida - 2, Protorthida - 1, Orthida - 5, Pentamerida - 1, Rhynchonellida - 4, Atrypida - 4, Athyridida - 3, Spiriferida - 4, Spiriferinida - 3) are described from the Szydłówek and Wietrznia Beds. Seven new species are introduced: Skenidioides cretus Halamski sp. nov., Biernatium minus Baliński sp. nov., Monelasmina montisjosephi Baliński sp. nov., Atryparia (Costatrypa) agricolae Halamski and Baliński sp. nov., Davidsonia enmerkaris Halamski sp. nov., Leptathyris gornensis Baliński sp. nov., and Echinocoelia parva Baliński sp. nov. Davidsonia enmerkaris Halamski sp. nov. is intermediate between Davidsonia Bouchard-Chantereaux, 1849 and Rugodavidsonia Copper, 1996 and is the youngest known representative of the suborder Davidsonioidea Copper, 1996. Skenidioides cretus Halamski sp. nov. is the last representative of the genus. Statistical investigation of a large sample of Spinatrypina (Exatrypa) explanata did not confirm the existence of two dimorphic

  13. Inferring seawater temperature over the past 2,500 years in the Southern California Bight on the basis of brachiopods (United States)

    Tomašových, Adam; Müller, Tamás; Kidwell, Susan M.


    Use of calcite δ18O in brachiopod shells in assessing past variations in seawater temperature remains poorly constrained in the absence of other methods due to vital effects and unknown variations in seawater density, salinity. Here, in order to evaluate past changes in seawater temperature of mainland shelf habitats off the Southern California Bight over the past 2,500 years, we analyze δ18O and Mg/Ca ratio of dead shells of the terebratulid brachiopod Laqueus erythraeus collected at 60-80 m water depths and age-dated by radiocarbon-calibrated amino acid racemization. These dead Holocene shells show excellent preservation (Mn concentrations < 10 ppm and Sr concentrations above 800 ppm). Although historical changes in sea-surface temperature in the southern California Bight were inferred on the basis of alkenones and δ18O in of planktonic foraminifers, temperature history of deeper shelf below storm wave base in this region remains unclear. First, we investigate thermal sensitivity of Mg/Ca ratio (using Laser Ablation Inductively Coupled Plasma Mass Spectrometry and wavelength-dispersive spectrometry) in the terebratulid brachiopod Laqueus erythraeus (collected in 1994 at Santa Catalina Island at 116 m water depth). At this depth, annual temperature range is relatively small (between 9-11°C), although at times of El Nino events in 1982-1983, 1986-1987, and 1992-1993, monthly temperature attained 13 °C. We find that δ18O measured along a growth profile of a shell precipitated in oxygen isotopic equilibrium with ambient seawater, and maxima in Mg/Ca ratio coincide with minima in δ18O, suggesting that fluctuations in Mg/Ca ratio trace temperature fluctuations, as observed also in other brachiopod species. Second, preliminary observations of Holocene shells show that Mg/Ca ratios show centennial-scale fluctuations but on average remain remarkably constant, with minima and maxima staying within intra-shell seasonal variations captured by extant specimens

  14. Prelude of benthic community collapse during the end-Permian mass extinction in siliciclastic offshore sub-basin: Brachiopod evidence from South China (United States)

    Wu, Huiting; He, Weihong; Weldon, Elizabeth A.


    Analysis of the Permian-Triassic palaeocommunities from basinal facies in South China provides an insight into the environmental deterioration occurring in the prelude to the mass extinction event. Quantitative and multivariate analyses on three brachiopod palaeocommunities from the Changhsingian to the earliest Triassic in basinal facies in South China have been undertaken in this study. Although the end-Permian extinction has been proved to be a one-stepped event, ecological warning signals appeared in the palaeocommunities long before the main pulse of the event. A brachiopod palaeocommunity turnover occurred in the upper part of the Clarkina changxingensis Zone, associated with a significant decrease of palaeocommunity diversity and brachiopod body size. During this turnover the dominant genera changed from Fusichonetes and Crurithyris (or/and Paracrurithyris) to the more competitive genus Crurithyris (or/and Paracrurithyris). The brachiopod palaeocommunity turnover was supposed to be triggered by the decreased marine primary productivity and increased volcanic activity. Moreover, such early warning signals are found not only in the deep-water siliceous facies, but also in the shallow-water clastic facies and carbonate rock facies in South China.

  15. Growth increments of the recent brachiopod Magellania venosa mechanically marked in Paso Comau and Comau Fjord, Chile, 2011/2012, supplement to: Baumgarten, Sebastian; Laudien, Jürgen; Jantzen, Carin; Häussermann, Verena; Försterra, Günter (2013): Population structure, growth and production of a recent brachiopod from the Chilean fjord region. Marine Ecology, 35(4), 401-413

    KAUST Repository

    Baumgarten, Sebastian


    Magellania venosa, the largest recent brachiopod, occurs in clusters and banks in population densities of up to 416 ind/m**2 in Comau Fjord, Northern Chilean fjord region. Below 15 m, it co-occurs with the mytilid Aulacomya atra and it dominates the benthic community below 20 m. To determine the question of why M. venosa is a successful competitor, the in situ growth rate of the brachiopod was studied and its overall growth performance compared with that of other brachiopods and mussels. The growth in length was measured between February 2011 and March 2012 after mechanical tagging and calcein staining. Settlement and juvenile growth were determined from recruitment tiles installed in 2009 and from subsequent photocensus. Growth of M. venosa is best described by the general von Bertalanffy growth function, with a maximum shell length (Linf) of 71.53 mm and a Brody growth constant (K) of 0.336/year. The overall growth performance (OGP index = 5.1) is the highest recorded for a rynchonelliform brachiopod and in the range of that for Mytilus chilensis (4.8-5.27), but lower than that of A. atra (5.74). The maximal individual production (PInd) is 0.29 g AFDM/ind/year at 42 mm shell length and annual production ranges from 1.28 to 89.25 g AFDM/year/m**2 (1-57% of that of A. atra in the respective fjords). The high shell growth rate of M. venosa, together with its high overall growth performance may explain the locally high population density of this brachiopod in Comau Fjord. However, the production per biomass of the population (P/B-ratio) is low (0.535) and M. venosa may play only a minor role in the food chain. Settling dynamics indicates that M. venosa is a pioneer species with low juvenile mortality. The coexistence of the brachiopod and bivalve suggests that brachiopod survival is affected by neither the presence of potential brachiopod predators nor that of space competitors (i.e. mytilids).

  16. Wolfcampian brachiopods from the Bird Spring Group, Wamp Spring area, Las Vegas Range, Clark County, Nevada ( USA). (United States)

    Mills, P.C.; Langenheim, R.L.


    The Wamp Spring section of the Bird Spring Group is subdivided into a basal platy limestone member, lower cliff-forming member, and upper cliff-forming member. Triticites, Schwagerina, and Schubertella kingi in the platy limestone member indicate an early Wolfcampian age. Schwagerina, Schubertella kingi, and a distinctive assemblage of brachiopods, similar to the West Texas fauna, indicate that the upper cliff-forming member is late Wolfcampian. The lower cliff-forming member is tentatively assigned to the middle Wolfcampian. The Wamp Spring sequence correlates temporally with the BSe 'formation' of the Bird Spring Group. The fossil-rich upper cliff-forming limestone member includes the new species Pontisia boodi, Crurithyris wampensis, and Calliprotonia(?) n. sp. A, as well as Hustedia culcitula, Crenispirifer(?) sp., Cenorhynchia(?) sp., Kutorginella(?) sp., marginiferids, lyssacine hexactinellid sponges, pleurotomarid and bellerophontid gastropods, cidaroid echinoids, rugose corals, cylindrical cryptostome bryozoans, and nuculids. -from Authors

  17. Boron isotopes in brachiopods during the end-Permian mass extinction: constraints on pH evolution and seawater chemistry (United States)

    Jurikova, Hana; Gutjahr, Marcus; Liebetrau, Volker; Brand, Uwe; Posenato, Renato; Garbelli, Claudio; Angiolini, Lucia; Eisenhauer, Anton


    The global biogeochemical cycling of carbon is fundamental for life on Earth with the ocean playing a key role as the largest and dynamically evolving CO2 reservoir. The boron isotope composition (commonly expressed in δ11B) of marine calcium carbonate is considered to be one of the most reliable paleo-pH proxies, potentially enabling us to reconstruct past ocean pH changes and understand carbon cycle perturbations along Earth's geological record (e.g. Foster et al., 2008; Clarkson et al., 2015). Brachiopods present an advantageous and largely underutilised archive for Phanerozoic carbon cycle reconstructions considering their high abundance in the geological record and its origin dating back to the early Cambrian. Moreover, their shell made of low-magnesium calcite makes these marine calcifiers more resistant to post-depositional diagenetic alteration of primary chemical signals. We have investigated the δ11B using MC-ICP-MS (Neptune Plus) and B/Ca and other elemental ratios (Mg/Ca, Sr/Ca, Al/Ca, Li/Ca, Ba/Ca, Na/Ca and Fe/Ca) using ICP-MS-Quadrupole (Agilent 7500cx) from the same specimens in pristine brachiopod shells from two sections from northern Italy during the Late Permian. These sections cover the δ13C excursion in excess of ˜4 ‰ (Brand et al., 2012) and are associated with major climate and environmental perturbations that lead to the mass extinction event at the Permian-Triassic boundary. Particular emphasis will be placed on the implications of our new paleo-pH estimates on the seawater chemistry during the Late Permian. Brand, U., Posenato, R., Came, R., Affek, H., Angiolini, L., Azmy, K. and Farabegoli, E.: The end-Permian mass extinction: A rapid volcanic CO2 and CH4-climatic catastrophe, Chemical Geology 323, 121-144, doi:10.1016/j.chemgeo.2012.06.015, 2012. Clarkson, M.O., Kasemann, S.A., Wood, R.A., Lenton, T.M., Daines, S.J., Richoz, S., Ohnemueller, F., Meixner, A., Poulton, S.W. and Tipper, E.T.: Ocean acidification and the Permo

  18. X-ray tomographic microscopy tightens affinity of the early Cambrian Oymurania to the brachiopod stem group

    Directory of Open Access Journals (Sweden)

    Artem Kouchinsky


    Full Text Available The geologically rapid biotic evolution in the early Cambrian is marked by the first appearance of major groups of animals in the fossil record (e.g., Budd and Jensen 2000; Kouchinsky et al. 2012. Along with the earliest crown-group representatives of the phylum Brachiopoda, more basal branches of the phylogenetic tree belonging to the stem-group Brachiopoda, such as tannuolinids and mikwitziids, diversified and became widespread during the early Cambrian (e.g., Williams and Holmer 2002; Balthasar et al. 2009; Skovsted et al. 2014. Synchrotron-radiation X-ray tomographic microscopy (SRXTM of Oymurania gravestocki reveals the microstructure of its calcium-phosphatic shell differentiated into two layers and intersecting systems of canals. The outer layer shows the acrotretoid columnar microstructure and the inner layer consists of continuous prismatic columns. Phosphatized setae preserved within tangential canals, as well as perpendicular canals of Micrina–Setatella type demonstrate homology with the tannuolinid Micrina and the mickwitziid Setatella. A unique and novel combination of microstructural features in Oymurania confirms its evolutionary position within the brachiopod stem group.

  19. Equatorial seawater temperatures and latitudinal temperature gradients during the Middle to Late Jurassic: the stable isotope record of brachiopods and oysters from Gebel Maghara, Egypt (United States)

    Alberti, Matthias; Fürsich, Franz T.; Abdelhady, Ahmed A.; Andersen, Nils


    The Jurassic climate has traditionally been described as equable, warmer than today, with weak latitudinal temperature gradients, and no polar glaciations. This view changed over the last decades with studies pointing to distinct climate fluctuations and the occasional presence of polar ice caps. Most of these temperature reconstructions are based on stable isotope analyses of fossil shells from Europe. Additional data from other parts of the world is slowly completing the picture. Gebel Maghara in the northern Sinai Peninsula of Egypt exposes a thick Jurassic succession. After a phase of terrestrial sedimentation in the Early Jurassic, marine conditions dominated since the end of the Aalenian. The stable isotope (δ18O, δ13C) composition of brachiopod and oyster shells was used to reconstruct seawater temperatures from the Bajocian to the Kimmeridgian at a palaeolatitude of ca. 3°N. Throughout this time interval, temperatures were comparatively constant aorund an average of 25.7°C. Slightly warmer conditions existed in the Early Bathonian ( 27.0°C), while the Kimmeridgian shows the lowest temperatures ( 24.3°C). The seasonality has been reconstructed with the help of high-resolution sampling of two oyster shells and was found to be very low (temperature gradients. During the Middle Jurassic, this gradient was much steeper than previously expected and comparable to today. During the Kimmeridgian, temperatures in Europe were generally warmer leading to weaker latitudinal gradients. Based on currently used estimates for the δ18O value of seawater during the Jurassic, reconstructed water temperatures for localities above the thermocline in Egypt and Europe were mostly lower than Recent sea-surface temperatures. These results improve our understanding of the Jurassic climate and its influence on marine faunal diversity patterns.

  20. Checklist of Recent thecideoid brachiopods from the Indian Ocean and Red Sea, with a description of a new species of Thecidellina from Europa Island and a re-description of T. blochmanni Dall from Christmas Island. (United States)

    Logan, Alan; Hoffmann, Jana; Lüter, Carsten


    Compilation of a checklist of Recent thecideoid brachiopods from the Indian Ocean and Red Sea indicates that members of this superfamily are represented by a small number of species. The subfamily Lacazellinae is represented by Ospreyella maldiviana from the Maldive Islands but the presence of Lacazella cannot yet be confirmed in the Indian Ocean as the holotype of Lacazella mauritiana from Mauritius is lost. The subfamily Thecidellininae is represented by Thecidellina blochmanni from Christmas Island in the eastern Indian Ocean and the Red Sea while a new species T. europa is here described from Europa Island in the Mozambique Channel. The subfamily Minutellinae is represented by Minutella minuta from Samper Bank and Walters Bank in the south-western Indian Ocean and in the Red Sea. Since the holotype of Thecidellina blochmanni from Flying Fish Cove, Christmas Island is also lost, this species is re-described and illustrated mainly from topotypes in the Museum für Naturkunde, Berlin, from which a suggested neotype has been selected.

  1. Obolus mrtvých a jeho význam pro datování archeologických pramenů

    Czech Academy of Sciences Publication Activity Database

    Bartošková, Andrea


    Roč. 38, č. 1 (2013), s. 129-138 ISSN 0231-5823 Institutional support: RVO:67985912 Keywords : Early Middle Ages * coins * graves * absolute dating * stratigraphy * pottery Subject RIV: AC - Archeology, Anthropology, Ethnology

  2. Geochemistry of the brachiopod Hemithiris psittacea from the Canadian Arctic

    DEFF Research Database (Denmark)

    Ullmann, Clemens V.; Korte, Christoph; Bitner, M. Aleksandra


    Fossil shell material from high palaeo-latitudes is increasingly studied for environmental forcings specific to these latitudes and as sensitive archives of climate change. However, more detailed studies of modern analogues are necessary for putting findings into global and high latitude perspect...

  3. Isotopic and elemental proxies in mollusc and brachiopod calcite

    DEFF Research Database (Denmark)

    Ullmann, Clemens Vinzenz

    composition of the oceans have changed. These changes of environmental conditions and seawater composition are primarily caused by plate tectonic processes, the cyclic variations of the Earth’s orbital parameters and living organisms, actively causing, and passively adapting to shifting conditions...

  4. Late Ordovician brachiopod distribution and ecospace partitioning in the Tvären crater system, Sweden

    DEFF Research Database (Denmark)

    Frisk, Åsa M.; Harper, David Alexander Taylor


    of the guild structure of the fauna permits explanation of a local biodiversity hotspot in otherwise low-diversity strata elsewhere in the Scandinavian region. The Tvaren impact event had an important palaeobiologic effect upon the fossil record as it served as a local pump and reservoir for biodiversity....... Moreover the development of new community types and narrowly-defined niches helped further drive both a and p biodiversity during a critical phase of the Great Ordovician Biodiversification Event (C) 2012 Elsevier B.V. All rights reserved....

  5. Biogeographic and bathymetric determinants of brachiopod extinction and survival during the Late Ordovician mass extinction

    DEFF Research Database (Denmark)

    Finnegan, Seth; Mac Ørum Rasmussen, Christian; Harper, David A. T.


    –Early Silurian genus extinctions and evaluate which extinction drivers are best supported by the data. The first (latest Katian) pulse of the LOME preferentially affected genera restricted to deeper waters or to relatively narrow (less than 35°) palaeolatitudinal ranges. This pattern is only observed...... in the latest Katian, suggesting that it reflects drivers unique to this interval. Extinction of exclusively deeper-water genera implies that changes in water mass properties such as dissolved oxygen content played an important role. Extinction of genera with narrow latitudinal ranges suggests that interactions...... between shifting climate zones and palaeobiogeography may also have been important. We test the latter hypothesis by estimating whether each genus would have been able to track habitats within its thermal tolerance range during the greenhouse–icehouse climate transition. Models including these estimates...

  6. Synchrotron radiation X-ray tomographic microscopy (SRXTM) of brachiopod shell interiors for taxonomy: Preliminary report


    Motchurova-Dekova Neda; Harper David A.T.


    Synchrotron radiation X-ray tomographic microscopy (SRXTM) is a non-destructive technique for the investigation and visualization of the internal features of solid opaque objects, which allows reconstruction of a complete three-dimensional image of internal structures by recording of the differences in the effects on the passage of waves of energy reacting with those structures. Contrary to X-rays, produced in a conventional X-ray tube, the intense synchrot...

  7. Lower Devonian Brachiopods and Stratigraphy of North Palencia (Cantabrian Mountains, Spain)

    NARCIS (Netherlands)

    Binnekamp, J.G.


    A continuous sequence of Devonian sediments is exposed in the northern part of the province of Palencia (NW-Spain), on the southern slope of the Cantabrian Mountains. This study concerns the stratigraphy and paleontology of the Lower Devonian formations. At the base of the sequence a clastic

  8. Larval Myogenesis in the Articulate Brachiopod Argyrotheca cordata (Risso, 1826)

    DEFF Research Database (Denmark)

    Altenburger, Andreas; Wanninger, Andreas Wilhelm Georg


    Brachiopoda is a metazoan phylum with a fossil record dating back to the lower Cambrium and comprises over 12,000 extinct and 400 recent species. Despite some recent progress using mainly palaeontological and molecular datasets, the discussion concerning the phylogenetic position of Brachiopoda r...

  9. Strontium chemostratigraphy of carbonate sediments - Pilot study of Silurian and Devonian brachiopods from the Prague Basin

    Czech Academy of Sciences Publication Activity Database

    Janoušek, V.; Hladil, Jindřich; Frýda, J.; Slavík, Ladislav; Šmíd, J.


    Roč. 13, - (2001), s. 68-69 ISSN 1210-9606. [Meeting of the Czech Tectonics Studies Group /6./. Donovaly - Nízké Tatry, 03.05.2001-06.05.2001] R&D Projects: GA AV ČR KSK3012103 Institutional research plan: CEZ:AV0Z3013912 Keywords : strontium chemostratigraphy * aquafacies * Devonian Subject RIV: DB - Geology ; Mineralogy

  10. Systematics of some Lower and Middle Devonian spiriferid brachiopods from Gaspe with a revision of the superfamily Delthyridoidea

    DEFF Research Database (Denmark)

    Bizzarro, Martin; Lespérance, P.J.


    new species aye described: Howellella (Howellella) forillonensis, Brachyspirifer (Brachyspirifer) briseboisi, Paraspirifer desbiensi, Brevispirifer florentinus, and B. quebecensis, The occurrence of Brevispirifer species with Middle Devonian chonetaceans confirms the presence of marine Eifelian strata...

  11. Checklist of recent brachiopod species collected during the Terrasses and Exbodi cruises in the New Caledonian region, SW Pacific (United States)

    Bitner, Maria Aleksandra


    Abstract Twenty species belonging to 16 genera, i.e. Neoancistrocrania, Novocrania, Basiliola, Basiliolella, Ebiscothyris, Stenosarina, Kanakythyris, Xenobrochus, Terebratulina, Eucalathis, Fallax, Frenulina, Septicollarina, Campages, Annuloplatidia, and Thecidellina have been identified in the material collected during the Terrasses and Exbodi cruises in the New Caledonian region, SW Pacific. The species Basiliolella grayi (Woodward, 1855) and Fallax neocaledonensis Laurin, 1997 are the most common in the studied collection, while Eucalathis murrayi (Davidson, 1878) is reported for the first time from the New Caledonian region. PMID:26798235

  12. Growth increments of the recent brachiopod Magellania venosa mechanically marked in Paso Comau and Comau Fjord, Chile, 2011/2012, supplement to: Baumgarten, Sebastian; Laudien, Jürgen; Jantzen, Carin; Häussermann, Verena; Försterra, Günter (2013): Population structure, growth and production of a recent brachiopod from the Chilean fjord region. Marine Ecology, 35(4), 401-413

    KAUST Repository

    Baumgarten, Sebastian; Laudien, Jü rgen; Jantzen, Carin; Hä ussermann, Verena; Fö rsterra, Gü nter


    and in the range of that for Mytilus chilensis (4.8-5.27), but lower than that of A. atra (5.74). The maximal individual production (PInd) is 0.29 g AFDM/ind/year at 42 mm shell length and annual production ranges from 1.28 to 89.25 g AFDM/year/m**2 (1

  13. Interrogation of distributional data for the End Ordovician crisis interval

    DEFF Research Database (Denmark)

    Mac Ørum Rasmussen, Christian; Harper, David Alexander Taylor


    The uppermost Ordovician–lowermost Silurian (Upper Katian–Rhuddanian) is surveyed with respect to the a- and ß-diversity of rhynchonelliformean brachiopods. The survey is based on new collections as well as existing literature, compiled in a large, georeferenced database. The brachiopod faunas ar...

  14. Ciliary and mucus-net filter feeding, with special reference to fluid mechanical characteristics

    DEFF Research Database (Denmark)

    Jørgensen, C.B.; Kiørboe, Thomas; Møhlenberg, F.


    feeding characterized by processing of water at low pressures (.ltoreq. 1 mm H2O). Mechanisms of water processing and particle retention in brachiopods and bivalves are compared. Laminar flow of through-currents and surface-currents in brachiopods is consistent with the hypothesis of capture of suspended...

  15. Metamorphosis in Craniiformea revisited

    DEFF Research Database (Denmark)

    Altenburger, Andreas; Wanninger, Andreas; Holmer, Lars E.


    We revisited the brachiopod fold hypothesis and investigated metamorphosis in the craniiform brachiopod Novocrania anomala. Larval development is lecithotrophic and the dorsal (brachial) valve is secreted by dorsal epithelia. We found that the juvenile ventral valve, which consists only of a thin...... brachiopods during metamorphosis to cement their pedicle to the substrate. N. anomala is therefore not initially attached by a valve but by material corresponding to pedicle cuticle. This is different to previous descriptions, which had led to speculations about a folding event in the evolution of Brachiopoda...

  16. Middle to Late Ordovician faunal studies from central Australia and Tasmania during the Great Ordovician Biodiversification Event

    DEFF Research Database (Denmark)

    Jakobsen, Kristian Grube

    A profound transformation of the marine biosphere took place during the Great Ordovician Biodiversification Event (GOBE), recognized as the longest interval of sustained biodiversification in the history of life. Successions through the Middle Ordovician Stairway Sandstone of the Amadeus Basin......, but the diversity is generally relatively low. The bivalves are the most dominant and diverse group in the central Australian fauna, whereas the brachiopods are the most dominant and diverse group in the Tasmanian fauna. Thirteen trilobite, five brachiopod, four conodont and 22 mollusc taxa have been documented......, but the preservation and low number of specimens prevent formal definition of additional new taxa. Although highly endemic at species level, the investigated brachiopod, trilobite and mollusc faunas display a high degree of overall similarity at generic level with Middle Ordovician faunas from North China. Other...

  17. Lenticellaria and Hillerella, new kraussinoid genera (Kraussinoidea, Brachiopoda) from Indo-Pacific and Red Sea waters: evolution in the subfamily Megerliinae. (United States)

    Simon, Eric G; Logan, Alan; Zuschin, Martin; Mainguy, Jerome; Mottequin, Bernard


    Two new kraussinid brachiopod genera, namely Lenticellaria gen. nov. and Hillerella gen. nov. are described from Pacific waters in the sub-equatorial zone in the Indonesian Archipelago, from Indian Ocean waters in Madagascar and from Red Sea waters in Egypt (Gulf of Aqaba) and Sudan. This fills the equatorial gap in the distribution of the superfamily Kraussinoidea, known from higher latitudes in both hemispheres. The micromorphic new material described is an excellent example of homeomorphy in brachiopods. It also provides new information on the distribution of the genus Megerlia sensu stricto and illustrates subtle variations in the evolutionary process of the reduced brachidium in Kraussinoidea.

  18. The lower Maastrichtian Hivdskud succession, Møns Klint, Denmark

    DEFF Research Database (Denmark)

    Jelby, M.E.; Thibault, Nicolas Rudolph; Surlyk, Finn


    with the cored boreholes Stevns-1 (Denmark) and ODP Site 762C (Indian Ocean). Hvidskud encompasses the 405 kyr eccentricity cycles Ma40513 – Ma40511 within magnetochron C31r. A sedimentation rate of 5.0 cm kyr-1 can be inferred from correlation to geochronological tie-points in ODP 762C, suggesting an age of ~70...... Maastrichtian in north-western Europe. Information on palaeo-seawater temperatures can be drawn from oxygen isotope records obtained from bulk rock samples and 24 micromorphic brachiopod specimens (Terebratulina faujasii). The brachiopod data show a clear diagenetic trend but point to an upper range...

  19. Quaternary deposits and weathered bedrock material as a source of dangerous radon emissions in Estonia

    Directory of Open Access Journals (Sweden)

    Petersell Valter


    Full Text Available The risk of dangerous radon emissions in Estonia is high, being among the highest in Europe. In almost 33 per cent of Estonian land area, the content of radon in soil-contained air exceeds the safe limit for unrestricted construction (50 kBq/m3. In such high radon-risk areas the concentration of radon in soil-contained air ranges from 50 to 400 kBq/m3, in a few cases reaching up to 2,100 kBq/m3 exceeding the permitted level for residential areas. The situation is particularly serious in the northernmost part of the country, where uranium-rich graptolite argillite (Dictyonema shale and the Obolus phosphorite are close to ground surface and their particles are constituent parts of Quaternary deposits. Radon emissions from bedrock have been investigated in detail, but to date Quaternary strata as a source of radon emissions are poorly studied. According to our measurements the highest concentrations of radon are related to tills containing clasts and fines of graptolite argillite and phosphorite. Glacial deposits include also granitoidal material, containing U, Th and K, which have been transported by glaciers from the outcrop areas of crystalline basement rocks in Finland and the Gulf of Finland. Due to weathering, outwash and repeated redeposition other genetic types are poorer in radioactive elements and they are weaker sources of radon.

  20. Late Devonian (Frasnian) bivalves from the Nocedo Formation: the results of Wilhelm Kegel’s 1927 field trip to northern Spain

    NARCIS (Netherlands)

    Amler, M.R.W.


    During a field trip to the Peña-Corada Unit of the southernmost Esla region of the Cantabrian Mountains in 1927, the German stratigrapher Wilhelm Kegel sampled brachiopods and bivalves from a section in the Laoz valley near La Ercina. The stratigraphic position is believed to be part of the Nocedo

  1. A functional collapse of persistent shell-gravel benthic ecosystem on the California shelf within the last century (United States)

    Tomasovych, Adam; Kidwell, Susan M.


    Death assemblages sampled from the muddy seabed of the inner and middle mainland Southern California continental shelf frequently contain dead shells of epifaunal terebratulid brachiopod and large-bodied scallop species that have not been encountered alive during annual surveys of this area over the last four decades. Instead, live-collected shelly benthos is dominated by infaunal species, especially chemosynthetic and deposit-feeding bivalves. Postmortem age-frequency distributions based on 190 individuals of the brachiopod Laqueus show (1) a mode between 100 and 300 years, (2) the absence of shells younger than 100 years old, and (3) the continuous presence of shells older than 300 years, ranging up to six thousands of years old, implying the relatively continuous active production of shells by this brachiopod species over millennia. The localized occurrence of small living populations of this brachiopod and of the scallops Chlamys and Euvola under the reduced sedimentation conditions along the outermost edge of the mainland shelf, and their occurrence on the sandy shelves of the isolated, offshore Channel Islands less affected by natural and anthropogenic runoff, indicates that, up until the last century, the inner and middle mainland shelf had also been characterized by extensive areas of mud-free, shell-gravel habitat. The shift in community structure to the spatially pervasive, infauna-dominated muddy habitats encountered today implies a change to higher siltation and sediment loading due to increased land clearance within recent centuries.

  2. Invertebrates and Plants :: NOAA Fisheries (United States)

    Invertebrates & Plants Species of Concern Threatened & Endangered Health & Stranding Marine Mammals Chart Partnerships Fisheries Home » Protected Resources » Species Invertebrates and Plants White Marine Sanctuary Mollusks, corals, and brachiopods are three groups of marine invertebrates. To view ESA

  3. Semiplanus (Productida, Brachiopoda) from the Carboniferous limestone of Kotaki, Niigata Prefecture, central Japan


    Ibaraki, Yousuke; Sato, Kiichi


    An Early Carboniferous large–sized productid brachiopod species, Semiplanus semiplanus (Schwetzow, 1922), is described from the Tsuchikurazawa Limestone (upper Visean–Serpukhovian), a limestone block within a Permian accretionary complex, distributed in the Kotaki area, Itoigawa City, Niigata Prefecture, central Japan. This is the first record of Semipanus species from Japan.

  4. Lipid and fatty acid fractions in Lingula anatina (Brachiopoda): an intertidal benthic fauna in the West Bengal-Orissa coast, India


    Samaresh Samanta; Tapas Kumar Das; Amalesh Choudhury; Susanta Kumar Chakraborty


    Objective: To record the fractional components of lipid and polyunsaturated fatty acids of Lingula anatina (L. anatina), a Precambrian intertidal benthic brachiopod, giving emphasis on -ω series group especially eicosapentaenoic acid (EPA) and docosahexaenoic acid (DHA) alongside assessing their biotransformation within the population and mangrove-estuarine associated community. Methods: Different biological samples after being collected from three contrasting study sites viz. ...

  5. Brachiopodové fauny triasu a jury v oblasti Salzkammergut v Rakousku

    Czech Academy of Sciences Publication Activity Database

    Siblík, Miloš


    Roč. 2006, - (2007), s. 166-167 ISSN 0514-8057 Institutional research plan: CEZ:AV0Z30130516 Keywords : brachiopods * Triassic * Jurassic Subject RIV: EG - Zoology

  6. An early 'handwritten' shell exchange catalogue of the Leiden Museum, with notes on the collectors Rethaan Macaré and Tischbein

    NARCIS (Netherlands)

    Bruggen, van A.C.


    A ‘handwritten’ (lithographed) shell exchange catalogue of the National Museum of Natural History, Leiden (The Netherlands), dated 1850 and containing 864 nominal taxa (861 molluscs and 3 brachiopods), is discussed in the context of the history of the museum. This catalogue must have been

  7. The highest-latitude Foliomena Fauna (Upper Ordovician, Portugal) and its palaeogeographical and palaeoecological significance

    DEFF Research Database (Denmark)

    Colmenar Lallena, Jorge; Pereira, Sofia; Sá, Artur Abreu


    of the Portuguese Buçaco-Penacova region (Central Iberian Zone). The three studied localities have yielded different associations, composed of 15 species of brachiopods and two trilobites, including both key elements of the deep water Foliomena Fauna and taxa typically from shallower benthic biotas, e...

  8. Taphonomy of a thick Terebratula bioherm from the Pliocene of southeastern Spain (United States)

    García-Ramos, Diego A.


    Brachiopods were extremely abundant during the Paleozoic era but underwent a dramatic loss of biodiversity at the Permo-Triassic boundary. The comparison of brachiopod and bivalve diversity through geological time shows that the latter were the most successful counterpart at best recovering from mass extinction events. Nonetheless, there are cases where Post-Paleozoic brachiopods stand out as the dominant marine benthos in particular environments, forming paucispecific brachiopod-dominated bioherms. This note describes an example of shallow-water brachiopod bioherm dominated by the terebratulid Terebratula calabra. The shell bed is found in mixed siliciclastic-temperate carbonate deposits of late Early Pliocene age nearby Águilas (southeastern Spain). This unique brachiopod concentration may be helpful to understand the particular success of large-sized brachiopods like Terebratula in Cenozoic environments typically dominated by bivalves. The bioherm attains 1.5 meters in thickness and crops out along a band up to 140 meters wide. The lithology consists of bioturbated fine-grained sands containing poorly sorted bioclasts, mostly fragments of Terebratula. This shell bed also records a diverse fauna, including five brachiopod genera, pectinids (4 genera), oysters (3 genera), in addition to rare gastropods, echinoids, bryozoans, etc. The density and sorting of bioclasts is laterally variable, and the biofabrics range from loosely dispersed to densely-packed, including examples of concave-up vertical stacking and nesting of shells. Most of the fragments of Terebratula preserve the posterior part of the shell only. These fragments generally display corrasion (rounded fractured margins, rounded to completely missing symphytium), bioerosion (prevailing the ichnogenera Entobia, Gnathichnus and Podichnus) and encrustation (mainly by bryozoans, Ancistrocrania, and Pododesmus). The good preservation of Pododesmus contrasts with that of most fragments of Terebratula, although

  9. Contributions to the Study of the Beginnings of the Orthodox Church in Western Rus’

    Directory of Open Access Journals (Sweden)

    Radosław Liwoch


    Full Text Available In 1882 T. Ziemięcki conducted in Pìdgìrcì (Lviv district, Ukraine a research excavation of two neighboring burial mounds which he called the Great Twin Barrows No. 1 and No. 2. In barrow No., he discovered on richly furnished double burial (probably of a married couple where he found two identical silver crosses. The crosses belong to the so called Scandinavian type (variant B.1 type 1.4.3 acc. J. Staecker. In barrow No. 2, he found a skeleton of a male buried with furnishings, including two crosses. The bigger one (lost was a bronze encolpion depicting the Crucifixion and the Virgin Orans, which can find analogies in the Danube countries. The smaller one is a silver St. Peter’s cross. In the mouths of the corpses found in both barrows there were tiny gold objects which could be interpreted as a local variation of the obolus of the dead custom. Both graves are a part of necropolis with druzhina burials. They date back to the end of the 10th century or the first half of the 11th century. The objects presented here are the earliest evidence to confirm the presence of a Christianized (Greek version of Christianity elite in the region of the Upper Bug and Upper Dniester, which is not burdened with chronological or interpretative reservation. The deceased were not members of the clergy but, nevertheless, representatives of the new official religion in a pagan Slavic population whose conversion had barely begun. It seems, therefore, that their presence in Pìdgìrcì can be considered the beginning of the Orthodox Church in the western part of the former Kievan Rus’.

  10. Homeobox gene expression in Brachiopoda

    DEFF Research Database (Denmark)

    Altenburger, Andreas; Martinez, Pedro; Wanninger, Andreas


    (ectoderm) specification with co-opted functions in notochord formation in chordates and left/right determination in ambulacrarians and vertebrates. The caudal ortholog, TtrCdx, is first expressed in the ectoderm of the gastrulating embryo in the posterior region of the blastopore. Its expression stays......The molecular control that underlies brachiopod ontogeny is largely unknown. In order to contribute to this issue we analyzed the expression pattern of two homeobox containing genes, Not and Cdx, during development of the rhynchonelliform (i.e., articulate) brachiopod Terebratalia transversa...... completion of larval development, which is marked by a three-lobed body with larval setae. Expression starts at gastrulation in two areas lateral to the blastopore and subsequently extends over the animal pole of the gastrula. With elongation of the gastrula, expression at the animal pole narrows to a small...

  11. Stratigraphy and macrofauna of the Lower Jurassic (Toarcian) Marrat Formation, central Saudi Arabia (United States)

    El-Sorogy, Abdelbaset S.; Gameil, Mohamed; Youssef, Mohamed; Al-Kahtany, Khaled M.


    The stratigraphy and macrofaunal content of the Lower Jurassic (Toarcian) Marrat Formation was studied at Khashm adh Dhibi, central Saudi Arabia. The studied succession is dominated by limestones and dolomites, with subordinate occurrences of sandstones, siltstones and claystones. The formation is highly fossiliferous with brachiopods, gastropods, bivalves, ammonites and echinoids, particularly the lower and upper members. Twenty nine species are identified, they include 7 species of brachiopods, 8 gastropods, 8 bivalves, 4 ammonites and 2 echinoids. Many of the identified fauna are correlated with Jurassic equivalents in Jordan, Italy, Morocco, Egypt and India. Three gastropod species: Globularia subumbilicata, Ampullospira sp., Purpuroidea peristriata and seven bivalve species: Palaeonucula lateralis, Chlamys (Radulopecten) fibrosa, Eligmus weiri, E.integer, E. asiaticus, Musculus somaliensis and Pholadomya orientalis were recognized for the first time in the Lower Jurassic deposits of Saudi Arabia.

  12. Hábitos de Vida da Associação “Schuchertella”agassizi – Pthychopteria eschwegei, Formação Maecuru,Devoniano, Bacia do Amazonas, Brasil.

    Directory of Open Access Journals (Sweden)

    Luiza Corral Martins de Oliveira Ponciano


    Full Text Available The fossils studied came from outcrops of the Maecuru and Curuá rivers, State of Pará, belonging to the upper strata of theMaecuru Formation, Eifelian age. In this formation, two associations of marine benthic invertebrate can be identified, each inferring adistinct paleoenvironment. The association analyzed occurs in medium and coarse sandstones and shows “Schuchertella” agassizi andPtychopeteria eschwegei as predominant organisms. It was idnetified 12 species of brachiopods, 20 species of bivalves, 12 of trilobites,8 of gastropods, 6 of bellerophonts, 3 of crinoids and 3 of tentaculitids in this association. Brachiopods are the most abundant organisms,responsible for approximately 50% of it, followed by tentaculitids and bivalves. Trilobites, gastropods and bellerophonts sum less than10% of the aforementioned association. Crinoids were not included in the quantitative analysis because they are only found as calicinalplates and disarticulated column disks. The brachiopods showed, through their morphofunctional characteristics, suspension feedersrecumbent (free-lying epifaunal life habit or a pedicle attached mode of life. The bivalves showed a predominant suspension feeder semiinfaunalhabit, attached by byssus threads to the substratum, and an infaunal mode of life. All the trilobites showed a predator/scavengerand vagile epifaunal habit. The epifaunal platyceratids represents the gastropods with low mobility (coprophagous/suspension feeders.The bellerophonts showed an epifaunal highly/medium mobility and grazing/predator habit. The tentaculitids having a semi-infaunal,suspension feeder habit. The suspension feeder forms (brachiopods, bivalves and tentaculitids account for over 90% of the organisms, withthe remaining percentage distributed among the predator/scavenger (trilobites, coprophagous/suspension feeders (gastropods, grazers/predators (bellerophonts and deposit feeders (rare bivalves. The predominance of suspension feeder forms

  13. Facies and age of the Oso Ridge Member (new), Abo Formation, Zuni Mountains, New Mexico (United States)

    Armstrong, A.K.; Stamm, R.G.; Kottlowski, F.E.; Mamet, B.L.; Dutro, J.T.; Weary, D.J.


    The Oso Ridge Member (new), at the base of the Abo Formation, nonconformably overlies Proterozoic rocks. The member consists of some 9m of conglomerate and arkose composed principally of fragments of the underlying Proterozoic metamorphic rocks; thin, fossiliferous limestone lenses are interbedded with the arkose. Biota from the lenses include a phylloid alga, foraminifers, conodonts, brachiopods, and molluscs. The age of the Oso Ridge Member is Virgilian Late Pennsylvanian) to Wolfcampian (Early Permian). -from Authors

  14. Revison of an Atheloptic Visean Trilobite Association in the Moravian Karst (Czech Republic

    Directory of Open Access Journals (Sweden)

    Rak Štěpán


    Full Text Available Recently two new trilobite occurrences were discovered during geological mapping of fossiliferous shales of the Březina Formation around the village of Březina in the Moravian Karst (Czech Republic. The newly discovered localities are extraordinary because of the unexpected occurrence of articulated trilobite exoskeletons associated with brachiopods including supporting spines. The new atheloptic association of Mississippian trilobites is described for the first time from the Moravian Karst.

  15. Additional Gigantoproductus species from the upper Visean-Namurian limestone of Kotaki, central Japan


    Ibaraki, Yousuke; Tazawa, Jun-ichi; Nakamura, Yukio


    A large productid brachiopod species, Gigantoproductus aurita (Bolkhovitinova, 1938), is described from the Tsuchikurazawa Limestone (upper Visean-Namurian), a large limestone block within a Permian accretionary complex of Kotaki, Itoigawa City, Niigata Prefecture, central Japan. This is the fourth described Gigantoproductus species from the Tsuchikurazawa Limestone. The range of G. aurita may extend to late Visean on the basis of the previous fossil records of the limestone.

  16. Prolonged Permian–Triassic ecological crisis recorded by molluscan dominance in Late Permian offshore assemblages


    Clapham, Matthew E.; Bottjer, David J.


    The end-Permian mass extinction was the largest biotic crisis in the history of animal life, eliminating as many as 95% of all species and dramatically altering the ecological structure of marine communities. Although the causes of this pronounced ecosystem shift have been widely debated, the broad consensus based on inferences from global taxonomic diversity patterns suggests that the shift from abundant brachiopods to dominant molluscs was abrupt and largely driven by the catastrophic effec...

  17. An early (1850) 'handwritten' shell exchange catalogue of the Leiden Museum, with notes on the collectors Rethaan Macaré and Tischbein


    Bruggen, van, A.C.


    A ‘handwritten’ (lithographed) shell exchange catalogue of the National Museum of Natural History, Leiden (The Netherlands), dated 1850 and containing 864 nominal taxa (861 molluscs and 3 brachiopods), is discussed in the context of the history of the museum. This catalogue must have been distributed in a limited edition to shell collectors interested in the exchange of specimens. Among the exchange partners of the museum were the well-known 19th century Dutch collector Lady (later Dowager) F...

  18. A new species of Conchicolites (Cornulitida, Tentaculita from the Wenlock of Gotland, Sweden

    Directory of Open Access Journals (Sweden)

    Olev Vinn


    Full Text Available A new cornulitid species, Conchicolites crispisulcans sp. nov., is described from the Wenlock of Gotland, Sweden. The undulating edge of C. crispisulcans sp. nov. peristomes is unique among the species of Conchicolites. This undulating peristome edge may reflect the position of setae at the tube aperture. The presence of the undulating peristome edge supports the hypothesis that cornulitids had setae and were probably related to brachiopods.

  19. Position of the Upper Devonian Frasnian--Famennian boundary in the central Appalachians

    Energy Technology Data Exchange (ETDEWEB)

    Rossbach, T.J. (Univ. of North Carolina, Chapel Hill, NC (United States). Dept. of Geology)


    Biostratigraphic analysis of eight Upper Devonian sections in VA and WV reveals that the section at Huttonsville, Randolph County, WV, is a key locality for determining the Frasnian-Famennian boundary. The Foreknobs Formation at Huttonsville indicates a higher stratigraphic placement of the Frasnian-Famennian boundary than has been generally assumed. Conodonts are not known within that section, so placement of the boundary uses the last occurrence of tentaculitids and the last and first occurrences of several species of brachiopods. It is believed that the Frasnian-Famennian boundary can be identified independently of the atrypoid brachiopods. Stratigraphic ranges of the cricoconarid Tentaculites discusses and the brachiopod Tropidoleptus carinatus, both considered Frasnian marker fossils, indicate that the Frasnian extends well into the Red Lick Member of the Foreknobs Formation at Huttonsville, with T. carinatus occurring up to 70% of the stratigraphic thickness of the Red Lick. The Famennian marker fossils A. angelica and C. sulcifer are both found at Huttonsville above the last recorded occurrence of all the Frasnian marker fossils. To the northeast of Huttonsville the Frasnian-Famennian series boundary has been placed by other workers within or at the top of the Pound Member of the Foreknobs Formation. This discrepancy implies that either the Pound Member is diachronous or that to the northeast paleoecologic factors caused local disappearances of critical fossils before their extinction at Huttonsville.

  20. Oxygen-isotope trends and seawater temperature changes across the Late Cambrian Steptoean positive carbon-isotope excursion (SPICE event) (United States)

    Elrick, M.; Rieboldt, S.; Saltzman, M.; McKay, R.M.


    The globally recognized Late Cambrian Steptoean positive C-isotope excursion (SPICE) is characterized by a 3???-5??? positive ??13C shift spanning SPICE represents a widespread ocean anoxic event leading to enhanced burial/preservation of organic matter (Corg) and pyrite. We analyzed ??18O values of apatitic inarticulate brachiopods from three Upper Cambrian successions across Laurentia to evaluate paleotemperatures during the SPICE. ??18O values range from ~12.5??? to 16.5???. Estimated seawater temperatures associated with the SPICE are unreasonably warm, suggesting that the brachiopod ??18O values were altered during early diagenesis. Despite this, all three localities show similar trends with respect to the SPICE ??13C curve, suggesting that the brachiopod apatite preserves a record of relative ??18O and temperature changes. The trends include relatively high ??18O values at the onset of the SPICE, decreasing and lowest values during the main event, and an increase in values at the end of the event. The higher ??18O values during the global extinction at the onset of the SPICE suggests seawater cooling and supports earlier hypotheses of upwelling of cool waters onto the shallow shelf. Decreasing and low ??18O values coincident with the rising limb of the SPICE support the hypothesis that seawater warming and associated reduced thermohaline circulation rates contributed to decreased dissolved O2 concentrations, which enhanced the preservation/burial of Corg causing the positive ??13C shift. ?? 2011 Geological Society of America.


    Directory of Open Access Journals (Sweden)



    Full Text Available The Guadalupian succession of the Huqf area (Sultanate of Oman represents a mega-sequence comprising the fluvial terrigenous Gharif Formation and the overlying marine Khuff Formation. The Khuff Fm. is subdivided into four members and is composed of marls and bioclastic limestones. The Khuff Fm. yields a rich fauna of brachiopods, conodonts, foraminifers, bivalves, gastropods, ostracods and cephalopods. The brachiopod fauna of the Khuff Fm. includes strophomenids, productids, orthids and terebratulids. The associated conodont fauna includes Hindeodus excavatus Behnken, Merrilina sp., M. praedivergens Kozur & Mostler, and Sweetina n. sp. (systematic descriptions of conodonts are given in the Paleontological Appendix. Foraminifers are represented by species of Miliolina and Rotaliina. The Khuff Fm. is given a Wordian age, based on brachiopods and conodonts. The depositional environment of the Khuff Fm. of southeastern Oman corresponds to the outer shelf of a large carbonate platform covering most of the Arabian Platform. The Khuff Fm. is interpreted as a major transgressive-regressive cycle related to differential subsidence. 

  2. The Carboniferous of the Western Karakoram (Pakistan) (United States)

    Gaetani, M.; Zanchi, A.; Angiolini, L.; Olivini, G.; Sciunnach, D.; Brunton, H.; Nicora, A.; Mawson, R.


    The results of the study of the Carboniferous successions in the western part of the Northern Karakoram during three geological expeditions are summarized here. Rocks of that period are not uniformly preserved in the several thrust sheets forming the Northern Karakoram. In most of them only the basal part of the Carboniferous, up to the Visean, is preserved, whilst in the Karambar thrust sheet a more complete section—previously almost unknown—is preserved. Four new lithostratigraphic units, time-constrained by brachiopod and conodont biostratigraphy, are described, from bottom to top: (1) the Margach Formation: prevailing dark shales with subordinate fine subarkoses and quartzarenites, up to 300 m thick (mid-Famennian to middle Tournaisian); (2) the Ribat Formation: grey crinoidal limestones passing upwards to dark marly limestones and marls, at least 300 m thick (middle Tournaisian to Serpukhovian); (3) the Lupsuk Formation: subarkoses to feldspathic quartzarenites in thick beds, alternating with dark shales and siltstones, up to 400 m thick (Serpukhovian to uppermost Carboniferous); (4) within the Lupsuk Formation a local member, the Twin Valleys Member, up to 100 m thick, a bioclastic limestone intercalation of post-Moscovian age, is distinguished. The Carboniferous successions are invariably sealed by the arkoses to quartzarenites of the Gircha Formation, 133 m above the base of which, in the Karambar area, an Asselian brachiopod fauna was recovered. The Carboniferous succession is interpreted as recording the evolution of the passive margin of the Northern Karakoram Terrane, from early rifting stage in the Late Devonian to syn-rift events during the Late Carboniferous. The basal part of the Gircha Formation, of latest Carboniferous-earliest Permian age, is considered to have been deposited above a break-up unconformity, linked to the early drifting in the seaway bordering the Karakoram. In the palaeontological appendix the most significant brachiopod taxa

  3. Uranium laterite from Ipora/Amorinopolis region- Goias, Brazil

    International Nuclear Information System (INIS)

    Fernandes, S.M.


    The present study gives an account of the uranium bearing laterite in the district of Amorinopolis, GO. Emphasis is given in the study of its mineralogy and of the mineralization controls. The uranium mineralization is chiefly found within the arkosic sandstones at the base of the Devonian Ponta Grossa Formation. The ore is tabular and concordant with the bedding, the controls being simultaneously litho-stratigraphic and biochemical. Narrow permeable horizons of arkosic sandstone lie between impermeable shale an siltstone layers. Within the permeable horizon fossil remains (probably brachiopods) are replaced by uranium minerals. The oxidized iron minerals may have acted to insulate and preserve the secondary soluble uranium minerals. (author)

  4. Late Ordovician (post-Sardic) rifting branches in the North Gondwanan Montagne Noire and Mouthoumet massifs of southern France

    DEFF Research Database (Denmark)

    Javier Álvaro, J.; Colmenar Lallena, Jorge; Monceret, Eric


    , and the subsequent Middle-Ordovician stratigraphic gap is related to the Sardic phase. Upper Ordovician sedimentation started in the rifting branches of Cabrières and Mouthoumet with deposition of basaltic lava flows and lahar deposits (Roque de Bandies and Villerouge formations) of continental tholeiite signature...... (CT), indicative of continental fracturing.The infill of both rifting branches followed with the onset of (1) Katian (Ka1-Ka2) conglomerates and sandstones (Glauzy and Gascagne formations), which have yielded a new brachiopod assemblage representative of the Svobodaina havliceki Community; (2) Katian...

  5. Precisely locating the Ordovician equator in Laurentia

    DEFF Research Database (Denmark)

    Jin, Jisuo; Harper, David A. T.; Cocks, L. Robin M.


    The Late Ordovician equatorial zone, like the zone today, had few hurricane-grade storms within 100 of the equator, as emphasized by the preservation of massive-bedded Thalassinoides ichnofacies in a trans-Laurentian belt more than 6000 km long, from the southwestern United States to North...... Greenland. That belt also includes nonamalgamated shell beds dominated by the brachiopod Proconchidium, which would not have been preserved after hurricane-grade storms. The belt lacks such storm-related sedimentary features as rip-up clasts, hummocky cross-stratification, or large channels. In contrast...

  6. Jurassic animals and algae in the flooring of Our Lady of Sorrows Church in Poznań

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    Antczak Mateusz


    Full Text Available The flooring of Our Lady of Sorrows Church in Poznań is made of Jurassic rocks from the Świętokrzyskie Mountains (also known as Holy Cross Mountains and contain abundant marine invertebrate fossils: sponges, bivalves, brachiopods, various families of cephalopods, etc. Some of them can be identified to the genus level. The fossils make it possible to describe the environment and ecosystem of the Jurassic sea and biostratigraphy of the sediment. There are also some significant inorganic structures, which suggest post-diagenetic tectonic movements.

  7. Limits in the evolution of biological form: a theoretical morphologic perspective. (United States)

    McGhee, George R


    Limits in the evolution of biological form can be empirically demonstrated by using theoretical morphospace analyses, and actual analytic examples are given for univalved ammonoid shell form, bivalved brachiopod shell form and helical bryozoan colony form. Limits in the evolution of form in these animal groups can be shown to be due to functional and developmental constraints on possible evolutionary trajectories in morphospace. Future evolutionary-limit research is needed to analyse the possible existence of temporal constraint in the evolution of biological form on Earth, and in the search for the possible existence of functional alien life forms on Titan and Triton that are developmentally impossible for Earth life.

  8. Early to Middle Jurassic palaeoenvironmental changes: High resolution δ13C and δ18O records from the UK

    DEFF Research Database (Denmark)

    Korte, Christoph; Hesselbo, Stephen; Ullmann, Clemens Vinzenz

    Low-Mg-calcite fossils, such as bivalves, belemnites and brachiopods, and bulk rocks have been extensively utilized to reconstruct past seawater chemistry and paleoenviron¬mental changes. Recent work on major bioevents demonstrated that particularly higher resolution stable isotope records...... are necessary to reveal short-term paleoenviron¬mental fluctuations and, in addition, to discover its causes. Here we present a new high resolution carbon and oxygen isotope dataset generated from low-Mg-calcite fossils, fossil wood and bulk rocks collected from Early to Middle Jurassic marine successions...

  9. Palaeoenvironmental changes in the Late Triassic (Rhaetian) of the Northern Calcareous Alps

    DEFF Research Database (Denmark)

    Mette, Wolfgang; Elsler, Armin; Korte, Christoph


    in the Kössen Formation and determine their significance for environmental and climatic conditions in the Rhaetian sea of the Eiberg Basin (Northern Calcareous Alps). For this purpose 60 δ 18O measurements on 43 articulate brachiopods samples from 8 different horizons were carried out. The results indicate...... to the lowest unit 3 of the Eiberg Member that parallels oxygen isotopes. This positive δ 13C trend is interrupted by a sudden ~1.5‰ negative excursion in the late Rhaetian (Late Rhaetian Event), a time span when the oxygen isotopes remain heavy....

  10. A lower Devonian point in the geological timescale

    International Nuclear Information System (INIS)

    Richards, J.R.; Barkas, J.P.; Vallance, T.G.


    The Gocup Granite, in southeastern New South Ales, intrudes low-grade metasediments tentatively assigned to the Ludlovian on fragmentary conodont evidence, and is overlain by a unit rich in volcanic material but containing a good brachiopod fauna of lower to middle Siegenian age. The K-Ar age of four muscovite samples is 409 +- 3 m.y.; a Rb-Sr whole-rock isochron on five samples yields a Model age 402 +- 3 m.y., initial 87 Sr/ 86 Sr 0.7090 +- 0.003. Preliminary data by X-ray fluorescence and unspiked strontium mass spectrometry illustrate the effects of some sources of experimental error. (auth.)

  11. Microfossils in the Ordovician erratic boulders from South-western Finland

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    Nõlvak, J.


    Full Text Available Chitinozoans, ostracods and acritarchs found in four glacially transported limestone boulders from the south-western coast of Finland have been studied in order to test the usefulness of these microfossil groups in age determinations. Also rare specimens of conodonts, inarticulated brachiopods and foraminifers were found. Baltic limestone (or Östersjö limestone was the most problematic, because only fossils with calcitic or phosphatic shells are preserved. It is concluded that the boulders identified correlate with the Uhaku and Rakvere stages of the Middle Ordovician.

  12. Tabasosphaera pustulosa nov. gen., nov. sp., a microproblematicum (foraminifer? from the Upper Triassic (Norian-Rhaetian reef limestone of Nayban Formation (NE Iran

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    Baba Senowbari-Daryan


    Full Text Available A microproblematicum with calcaerous test, Tabasosphaera pustulosa nov. gen., nov. sp., a hollow spherical microfossil with pustule-like elements on the outer surface, is described from Upper Triassic (Norian-Rhaetian reef limestones exposed near the town of Ali-Abad, south of Tabas (northeast Iran. Tabasosphaera is an endobiotic organism living within the cavities of other organisms, such as brachiopods, worm tubes, or in sponge spongocoels. The systematic position of Tabasosphaera, its interspecific association and the fossil assemblage of which it makes part is discussed. Most probably Tabasosphaera represents a foraminifer belonging to the group Lagenina.

  13. Stratigraphy and facies development of the marine Late Devonian near the Boulongour Reservoir, northwest Xinjiang, China (United States)

    Suttner, Thomas J.; Kido, Erika; Chen, Xiuqin; Mawson, Ruth; Waters, Johnny A.; Frýda, Jiří; Mathieson, David; Molloy, Peter D.; Pickett, John; Webster, Gary D.; Frýdová, Barbora


    Late Devonian to Early Carboniferous stratigraphic units within the 'Zhulumute' Formation, Hongguleleng Formation (stratotype), 'Hebukehe' Formation and the Heishantou Formation near the Boulongour Reservoir in northwestern Xinjiang are fossil-rich. The Hongguleleng and 'Hebukehe' formations are biostratigraphically well constrained by microfossils from the latest Frasnian linguiformis to mid-Famennian trachytera conodont biozones. The Hongguleleng Formation (96.8 m) is characterized by bioclastic argillaceous limestones and marls (the dominant facies) intercalated with green spiculitic calcareous shales. It yields abundant and highly diverse faunas of bryozoans, brachiopods and crinoids with subordinate solitary rugose corals, ostracods, trilobites, conodonts and other fish teeth. The succeeding 'Hebukehe' Formation (95.7 m) consists of siltstones, mudstones, arenites and intervals of bioclastic limestone (e.g. 'Blastoid Hill') and cherts with radiolarians. A diverse ichnofauna, phacopid trilobites, echinoderms (crinoids and blastoids) together with brachiopods, ostracods, bryozoans and rare cephalopods have been collected from this interval. Analysis of geochemical data, microfacies and especially the distribution of marine organisms, which are not described in detail here, but used for facies analysis, indicate a deepening of the depositional environment at the Boulongour Reservoir section. Results presented here concern mainly the sedimentological and stratigraphical context of the investigated section. Additionally, one Late Devonian palaeo-oceanic and biotic event, the Upper Kellwasser Event is recognized near the section base.

  14. Microbialites in the shallow-water marine environments of the Holy Cross Mountains (Poland) in the aftermath of the Frasnian-Famennian biotic crisis (United States)

    Rakociński, Michał; Racki, Grzegorz


    Microbial carbonates, consisting of abundant girvanellid oncoids, are described from cephalopod-crinoid and crinoid-brachiopod coquinas (rudstones) occurring in the lowermost Famennian of the Holy Cross Mountains, Poland. A Girvanella-bearing horizon (consist with numerous girvanellid oncoids) has been recognised at the Psie Górki section, and represents the northern slope succession of the drowned Dyminy Reef. This occurrence of microbialites in the aftermath of the Frasnian-Famennian event is interpreted as the result of opportunistic cyanobacteria blooms, which, as 'disaster forms', colonised empty shallow-water ecological niches during the survival phase following the Frasnian metazoan reef collapse, due to collapsed activity of epifaunal, grazing, and/or burrowing animals. The anachronistic lithofacies at Psie Górki is linked with catastrophic mass mortality of the cephalopod and crinoid-brachiopod communities during the heavy storm events. This mass occurrence of girvanellid oncoids, along with Frutexites-like microbial shrubs and, at least partly, common micritisation of some skeletal grains, records an overall increase in microbial activity in eutrophic normal marine environments. Microbial communities in the Holy Cross Mountains are not very diverse, being mainly represented by girvanellid oncoids, and stand in contrast to the very rich microbial communities known from the Guilin area (China), Canning Basin (Australia) and the Timan-northern Ural area (Russia). The association from Poland is similar to more diverse microbial communities represented by oncoids, trombolites and stromatolites, well known from the Canadian Alberta basin.

  15. Hirnantia sagittifera (Brachiopoda and Mucronaspis mucronata s.l. (Trilobita in the Upper Ordovician of the East Baltic: taxonomy and distribution

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    Linda Hints


    Full Text Available The brachiopod Hirnantia sagittifera (M’Coy and trilobite morphs of the genus Mucronaspis from the topmost Ordovician Porkuni Stage of the central East Baltic are described and compared with those from the Hirnantian Stage of other regions. These important Hirnantian taxa occur in the Livonian Tongue of the Central Baltoscandian Facies Belt of the Baltic Basin, where the Porkuni Stage is represented by the non-graptolitic Kuldiga and Saldus formations. Hirnantia sagittifera appears in the lowermost part of the Porkuni Stage and is rather widely distributed in the basin in spite of its rare finds in each locality. Our study of trilobites of the genus Mucronaspis has enabled us to observe morphological changes in its exoskeleton in time and to identify a succession of five morphotypes (morphs. In some characteristics these morphs are similar to those of different alleged species of the genus Mucronaspis (M. olini, M. danai, M. ganabina, M. mucronata but they cannot be definitely assigned to any of these species due to some variances. However, here for the first time a stratigraphically ordered collection is presented, which deserves attention in revising the taxonomy of highly variable Mucronaspis. The described brachiopods and trilobites occur mainly in the strata correlated with the Normalograptus extraordinarius graptolite Biozone. However, the uppermost finds of both taxa come from strata correlated with the N. persculptus Biozone.

  16. Abiotic and biotic responses to Milankovitch-forced megamonsoon and glacial cycles recorded in South China at the end of the Late Paleozoic Ice Age (United States)

    Fang, Qiang; Wu, Huaichun; Hinnov, Linda A.; Tian, Wenqian; Wang, Xunlian; Yang, Tianshui; Li, Haiyan; Zhang, Shihong


    At the end of the Late Paleozoic Ice Age (LPIA) from late Early Permian to early Late Permian, the global climate was impacted by a prevailing megamonsoon and Gondwanan deglaciation. To better understand the abiotic and biotic responses to Milankovitch-forced climate changes during this time period, multi-element X-ray fluorescence (XRF) geochemistry analyses were conducted on 948 samples from the late Early-late Middle Permian Maokou Formation at Shangsi, South China. The Fe/Ti, S/Ti, Ba/Ti and Ca time series, which were calibrated with an existing "floating" astronomical time scale (ATS), show the entire suite of Milankovitch rhythms including 405 kyr long eccentricity, 128 and 95 kyr short eccentricity, 33 kyr obliquity and 20 kyr precession. Spectral coherency and cross-phase analysis reveals that chemical weathering (monitored by Fe/Ti) and upwelling (captured by S/Ti and Ba/Ti) are nearly antiphase in the precession band, which suggests a contrast between summer and winter monsoon intensities. Strong obliquity signal in the Ba/Ti series is proposed to derive from changes in thermohaline circulation intensity from glaciation dynamics in southern Gondwana. The abundance of foraminifer, brachiopod and ostracod faunas within the Maokou Formation were mainly controlled by the 1.1 Myr obliquity modulation cycle. The obliquity-forced high-nutrient and oxygen-depleted conditions generally produced a benthic foraminifer bloom, but threatened the brachiopod and ostracod faunas.

  17. Embryonic chirality and the evolution of spiralian left–right asymmetries (United States)


    The group Spiralia includes species with one of the most significant cases of left–right asymmetries in animals: the coiling of the shell of gastropod molluscs (snails). In this animal group, an early event of embryonic chirality controlled by cytoskeleton dynamics and the subsequent differential activation of the genes nodal and Pitx determine the left–right axis of snails, and thus the direction of coiling of the shell. Despite progressive advances in our understanding of left–right axis specification in molluscs, little is known about left–right development in other spiralian taxa. Here, we identify and characterize the expression of nodal and Pitx orthologues in three different spiralian animals—the brachiopod Novocrania anomala, the annelid Owenia fusiformis and the nemertean Lineus ruber—and demonstrate embryonic chirality in the biradial-cleaving spiralian embryo of the bryozoan Membranipora membranacea. We show asymmetric expression of nodal and Pitx in the brachiopod and annelid, respectively, and symmetric expression of Pitx in the nemertean. Our findings indicate that early embryonic chirality is widespread and independent of the cleavage programme in the Spiralia. Additionally, our study illuminates the evolution of nodal and Pitx signalling by demonstrating embryonic asymmetric expression in lineages without obvious adult left–right asymmetries. This article is part of the themed issue ‘Provocative questions in left–right asymmetry’. PMID:27821523

  18. Patterns of fossil distributions within their environmental context from the Middle Triassic in South Canyon, Central Nevada, USA

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    Pedro M. Monarrez


    Full Text Available The Middle Triassic records the return of diverse marine communities after the severe effects of the end-Permian mass extinction. This diversification leads to the Mesozoic/modern adaptive radiation resulting in substantial changes in marine communities in comparison to their Paleozoic predecessors. This analysis focuses on the faunal abundance, ecological patterns, and environmental interpretation of a Middle Triassic section in Central Nevada. Twelve bulk samples were collected. Visible fossils were identified and tallied from hand samples and thin-sections were used to aid in environmental interpretation. Beginning in the Late Anisian, we observed an ammonoid dominated to flat-clam, epifaunal dominated benthic community within a muddy, quiet, inner shelf depositional environment. Through time, epifaunal bivalves dominate within a middle shelf environment followed by an increase in infaunalization and shell-thickness. During this time the presence of oncoids and the reported finding of corals suggest the middle shelf environment gave way to a higher energy patch reef shelf edge environment. Finally, we observe epifaunal brachiopods communities at the top of our section deposited in a middle shelf environment. In sum, we observe the dominance of modern taxa (i.e., bivalves with Paleozoic ecologies (i.e., epifaunal, followed by the dominance of modern taxa with Modern ecologies (i.e., infaunal, thick shells and then a return to Paleozoic taxa (i.e., brachiopods and Paleozoic ecologies within an overall transgressive environment.

  19. Biofacies and habitats of Brereton Limestone member (Carbondale Formation, Middle Pennsylvanian), Southwestern Illinois

    Energy Technology Data Exchange (ETDEWEB)

    O' Connell, D.B.; Utgaard, J.


    The Brereton Limestone is a shallow-water, open-marine carbonate deposited over peat or delta-plain muds after delta abandonment and a marine transgression. Data on autecology, lithology, insoluble residue content, and thickness were used to interpret the habitats of each biofacies. Biofacies V, a low-diversity biofacies dominated by brachiopods and ostracods, occupied turbid-water, mud- or shelly mud-bottom areas during influxes of detrital clays late in the abandonment of the Herrin delta and, also, early in the construction of the Jamestown delta. Low-relief carbonate mud mounds accumulated within and around baffles provided by thickets of phylloid algae and foraminifers, are capped locally by biofacies VI, a low-diversity biofacies dominated by ostracods. Biofacies VI, occupied the high subtidal to supratidal crests of algal mud mounds which had a stressed (possibly hypersaline) environment. Deeper water mud mounds were occupied by either Biofacies III, a crinoid-mixed fossil biofacies, or by Biofacies IV, which is dominated by fusulinids, strophomenids, and trilobites. Biofacies II, dominated by sponges, mollusks, and impunctate brachiopods, generally occurred on the flanks of the shallow-water mounds. Biofacies I, III, and IV also occurred in broad, muddy intermound areas and Biofacies III in narrow, winnowed intermound areas.

  20. Mass extinctions: Ecological selectivity and primary production (United States)

    Rhodes, Melissa Clark; Thayer, Charles W.


    If mass extinctions were caused by reduced primary productivity, then extinctions should be concentrated among animals with starvation-susceptible feeding modes, active lifestyles, and high-energy budgets. The stratigraphic ranges (by stage) of 424 genera of bivalves and 309 genera of articulate brachiopods suggest that there was an unusual reduction of primary productivity at the Cretaceous/Tertiary (K/T) boundary extinction. For bivalves at the K/T, there were (1) selective extinction of suspension feeders and other susceptible trophic categories relative to deposit feeders and other resistant categories, and (2) among suspension feed-ers, selective extinction of bivalves with active locomotion. During the Permian-Triassic (P/Tr) extinction and Jurassic background time, extinction rates among suspension feeders were greater for articulate brachiopods than for bivalves. But during the K/T event, extinction rates of articulates and suspension-feeding bivalves equalized, possibly because the low-energy budgets of articulates gave them an advantage when food was scarce.

  1. Phylogeny and mitochondrial gene order variation in Lophotrochozoa in the light of new mitogenomic data from Nemertea

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    von Döhren Jörn


    Full Text Available Abstract Background The new animal phylogeny established several taxa which were not identified by morphological analyses, most prominently the Ecdysozoa (arthropods, roundworms, priapulids and others and Lophotrochozoa (molluscs, annelids, brachiopods and others. Lophotrochozoan interrelationships are under discussion, e.g. regarding the position of Nemertea (ribbon worms, which were discussed to be sister group to e.g. Mollusca, Brachiozoa or Platyhelminthes. Mitochondrial genomes contributed well with sequence data and gene order characters to the deep metazoan phylogeny debate. Results In this study we present the first complete mitochondrial genome record for a member of the Nemertea, Lineus viridis. Except two trnP and trnT, all genes are located on the same strand. While gene order is most similar to that of the brachiopod Terebratulina retusa, sequence based analyses of mitochondrial genes place nemerteans close to molluscs, phoronids and entoprocts without clear preference for one of these taxa as sister group. Conclusion Almost all recent analyses with large datasets show good support for a taxon comprising Annelida, Mollusca, Brachiopoda, Phoronida and Nemertea. But the relationships among these taxa vary between different studies. The analysis of gene order differences gives evidence for a multiple independent occurrence of a large inversion in the mitochondrial genome of Lophotrochozoa and a re-inversion of the same part in gastropods. We hypothesize that some regions of the genome have a higher chance for intramolecular recombination than others and gene order data have to be analysed carefully to detect convergent rearrangement events.

  2. Biogenic hardparts: Difficult archives of the geological past (Invited) (United States)

    Immenhauser, A.; Schone, B. R.; Hoffmann, R.; Niedermayr, A.


    Biomineralized exo- or endoskeletons of fossil marine invertebrates are widespread and diverse components of the Phanerozoic rock record of Earth's past and present oceans. Exoskeletons serve as protection against environmental pressure or predators, whilst endoskeletons can act as support or serve as an attachment for muscles and ligaments and hence as a mechanism for transmitting muscular forces. Biogenic hard parts represent sophisticated products resulting from the hierarchical interaction of inorganic minerals (95%) and macromolecular organic matrices, forming commonly less than 5%. The significance of many biogenic carbonate archives lies in the time-resolved growth patterns and their ability to record ambient environmental conditions in the form of multiple geochemical properties (multi-proxy archives) that have been widely used to assess past oceanic seawater properties. Here, we compile and review published work dealing with crystallization pathways of skeletal hard parts secreted by mollusks (i.e., bivalves and cephalopods) as well as brachiopods as widely used archives of ancient neritic epeiric settings. Bivalves and cephalopods (e.g., extinct ammonoids and belemnites and extant Sepia, Nautilus and Spirula) all form accretionary calcitic, aragonitic or vateritic skeletal hard parts. Despite the fact that mollusks and brachiopods form part of very different branches of the animal phylogenetic tree, their biomineralization strategies are surprisingly similar. Our main focus lies in a critical assessment of the complex pathways of ions and aquo-complexes from their source (seawater) to the final product (biomineral). We do this as an attempt to critically test the commonly held hypothesis that many fossil hard parts precipitated (under favorable conditions and pending subsequent diagenetic alteration) in equilibrium with seawater. Two main observations stand out: (1) the present knowledge on pathways and mechanisms (e.g., ion channel trans-membrane or

  3. Conodont succession and reassessment of major events around the Permian-Triassic boundary at the Selong Xishan section, southern Tibet, China (United States)

    Yuan, Dong-Xun; Zhang, Yi-Chun; Shen, Shu-Zhong


    A major discrepancy for the age of the Selong Group from middle Cisuralian (Early Permian) to Changhsingian resulted from previous reports of Sakmarian, Kungurian and Guadalupian (Middle Permian) conodonts and Lopingian (Late Permian) brachiopods. Recently, Cisuralian and Guadalupian conodonts were reported again from the Selong Group and the basal part of the Kangshare Formation at the Selong section, but the age discrepancy remains. We present our conodont materials based on large samples collected from the Selong Group and our interpretation based on identifications using a sample population approach. Three conodont zones are recognized in our re-investigation of the upper part of the Selong Group. They include the Vjalovognathus sp., the Mesogondolella hendersoni, and the M. sheni zones, in ascending order. These zones are overlain by the basal Triassic Hindeodus parvus Zone and the Otoceras woodwardi Zone. Our reassessment of conodonts reported by previous studies from Selong and nearby sections suggest that all specimens consistently point to a Lopingian age; the upper part of the Selong Group is latest Changhsingian in age based on the presence of Clarkina orchardi and Mesogondolella sheni. Previously reported early Cisuralian and Guadalupian conodonts are misidentified using a form species concept. A hiatus may be present at the erosional surface between the Selong Group and the Waagenites Bed of the basal part of the Kangshare Formation. However, the hiatus is minimal because conodont and brachiopod assemblages above and below this surface are very similar, and it results from a latest Changhsingian transgression just before the extinction that follows a global latest Changhsingian regression. There is a distinct rapid end-Permian mass extinction at Selong within the Waagenites Bed, as indicated by the disappearances of all benthic brachiopods, rugose corals and Permian bryozoans. The burst of Clarkina species in the Waagenites Bed and throughout the

  4. Cornulitids (tubeworms) from the Late Ordovician Hirnantia fauna of Morocco (United States)

    Gutiérrez-Marco, Juan Carlos; Vinn, Olev


    Two species of cornulitids, Cornulites gondwanensis sp. nov. and C. aff. shallochensis Reed are described from the Hirnantian of Morocco, within an assemblage representative of the Hirnantia brachiopod fauna occurring near the Ordovician South Pole. The dominance of aggregated and solitary free forms could be explained by particular sedimentary environments preceding the Hirnantian glaciation and the latest Ordovician Extinction Event. The diversity of cornulitids in the Late Ordovician of Gondwana and related terranes was relatively low, and less diverse than the cornulitids of Laurentia and Baltica. Hirnantian cornulitids from Morocco do not resemble Late Ordovician cornulitids of Baltica and Laurentia. Moroccan cornulitids seem to be closely allied to some older Gondwanan cornulitids, especially Sardinian ones. They resemble species described from the Late Ordovician and Llandovery of Scotland suggesting a palaeobiogeographic link.

  5. The mitochondrial genome of phoronis architecta--Comparisons demonstrate that phoronids are lophotrochozoan protostomes

    Energy Technology Data Exchange (ETDEWEB)

    Helfenbein, Kevin G.; Boore, Jeffrey L.


    The proper reconstruction of the relationships among the animal phyla is central to interpreting patterns of animal evolution from the genomic level to the morphological level. This is true not only of the more speciose phyla, but also of smaller groups. We report here the nearly complete DNA sequence of the mitochondrial genome of the phoronid Phoronis architecta, which has a gene arrangement remarkably similar to that of a protostome animal, the chiton Katharina tunicata. Evolutionary analysis of both gene arrangements and inferred amino acid sequences of these taxa, along with those of three brachiopods and other diverse animals, strongly supports the hypothesis that lophophorates are part of the large group that includes mollusks and annelids, i.e., the Lophotrochozoa, and solidly refutes the alternative of their being deuterostomes.

  6. Lophotrochozoan mitochondrial genomes

    Energy Technology Data Exchange (ETDEWEB)

    Valles, Yvonne; Boore, Jeffrey L.


    Progress in both molecular techniques and phylogeneticmethods has challenged many of the interpretations of traditionaltaxonomy. One example is in the recognition of the animal superphylumLophotrochozoa (annelids, mollusks, echiurans, platyhelminthes,brachiopods, and other phyla), although the relationships within thisgroup and the inclusion of some phyla remain uncertain. While much ofthis progress in phylogenetic reconstruction has been based on comparingsingle gene sequences, we are beginning to see the potential of comparinglarge-scale features of genomes, such as the relative order of genes.Even though tremendous progress is being made on the sequencedetermination of whole nuclear genomes, the dataset of choice forgenome-level characters for many animals across a broad taxonomic rangeremains mitochondrial genomes. We review here what is known aboutmitochondrial genomes of the lophotrochozoans and discuss the promisethat this dataset will enable insight into theirrelationships.


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    Full Text Available The bivalve Daonella Mojsisovics, 1874 is very common in the Middle Triassic pelagic facies, whereas the record of this genus from shallow water limestones is rare. In the present paper a new species of Daonella, named D. pseudograbensis, is described from the Esino Limestone, a Ladinian (Middle Triassic carbonate platform in the central Southern Alps. The species is described from Brembana Valley, where the Esino Limestone is rather rich in bioclastic lenses yielding faunas with bivalves, cephalopods, gastropods, brachiopods, corals and calcareous algae. Daonella pseudograbensis n. sp. is based on very well preserved specimens, which are often articulated and closed, all coming from the same locality. The new species shows a narrow range of intraspecific and ontogenetic morphologic variations. It is easy distinguishable from the other species of the genus for the outline and ornamentation; it therefore differs from D. grabensis Kittl, 1912, the most similar species, for the longer anterior dorsal margin.Pdf

  8. Aspectos gerais sobre a alimentação do cangulo, Balistes vetula Linnaeus, 1758 (Pisces - Balistidae no estado de Pernambuco - Brasil

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    Técio Luiz Macêdo Costa


    Full Text Available The Balistidae, especially Balistes vetula are commom fishes in tropical waters from Northeastern Brazil. Studies about these fishes captured at the coastal area of Recife between Pina; Boa Viagem, Piedade and Candeias beaches were carried out in order to determine the diet, and to verify a possible diet diversification between the sexes, and times of the year. Males and females had a food diet consisting mostly of molluscs belonging the classes Scaphopoda, Bivalvia, Gastropoda and crustaceans and fishes. Echinoderms, foraminifera, bryozoa, sponges, polychaete worms, coelenterates and brachiopods were also observed in smaller proportions. Both males and females had a carnivorous benthic diet. Growth and changes in the seasons were not reflected by changes in the diet.

  9. Subsurface example of a dolomitized middle Guadalupian (Permian) reef from west Texas

    Energy Technology Data Exchange (ETDEWEB)

    Longacre, S.A.


    A middle Guadalupian organic buildup was cored in the North McElroy Unit (NMU No. 3713 well) in Upton County, W. Texas. Fusulinid control indicates the buildup is equivalent to the Goat Seep reef that crops out in the Guadalupe Mt. The organic buildup at North McElroy consists of boundstones and associated flank grainstones. The reef biota is dominated by ramose and encrusting bryozoans, numerous types of calcareous sponges, the problematic encrusting organism Tubiphytes, encrusting algae, crinoids, brachiopods, and trilobites. Among the more significant aspects of diagenesis are mechanical degradation and biologic degradation, leaching, gypsum-anhydrite transformations, cementation, replacement of all calcite by dolomite, cementation and replacement by baroque dolomite, fracturing, and replacement of carbonate by anhydrite. 17 references.

  10. Sedimentary uranium deposit of the Ipora/Amorinopolis region, state of Goias, Brazil

    International Nuclear Information System (INIS)

    Fernandes, S.M.; Leonardos, O.H.


    The uranium mineralization is chiefly found within arkosic sandstones at the base of the Devonian Ponta Grossa Formation. The ore is tabular and concordant with the bedding, the controls being simultaneously litho-stratigraphical and biochemical. Narrow permeable horizons of arkosic sandstone lie between impermeable shale and siltstone layers. Within the permeable horizon, the fossil remains (probably brachiopods) are replaced by uranium minerals. The oxidized iron minerals may have acted as to insulate and preserve the secondary soluble uranium minerals. The mineral paragenesis is represented by renardite, meta - autunite I, fourmarierite, Koninckite, ranquilite, meta-uranocircite II, barite, apatite, calophane, wavelite, varscite, an unnamed uranium mineral, quartz, calcedony, goethite, lepidocrocite and hematite. (Author) [pt

  11. Combined oxygen- and carbon-isotope records through the Early Jurassic: multiple global events and two modes of carbon-cycle/temperature coupling

    DEFF Research Database (Denmark)

    Hesselbo, Stephen P.; Korte, Christoph


    , to the extent that meaningful comparisons between these events can begin to be made. Here we present new carbon and oxygen isotope data from mollusks (bivalves and belemnites) and brachiopods collected through the marine Early Jurassic succession of NE England, including the Sinemurian-Plienbachian boundary...... GSSP. All materials have been screened by chemical analysis and scanning electron microscopy to check for diagenetic alteration. Analysis of carbon isotopes from marine calcite is supplemented by analysis of carbon-isotope values from fossil wood collected through the same section. It is demonstrated...... that both long-term and short-term carbon-isotope shifts from the UK Early Jurassic represent global changes in carbon cycle balances. The Sinemurian-Pliensbachian boundary event is an event of global significance and shows several similarities to the Toarcian OAE (relative sea-level change, carbon-isotope...

  12. New data on internal morphology of exceptionally preserved Nannirhynchia pygmaea (Morris, 1847 from the Lusitanian Basin (Brachiopoda, Early Jurassic, Portugal

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    M. Schemm-Gregory


    Full Text Available Pyritized internal moulds of articulated shells of the Early Jurassic brachiopod taxon Nannirhynchia pygmaea were found in beds closely below the early Toarcian oceanic anoxic event in the Polymorphum Zone in Portugal. The material allows a detailed study of the outline of the muscle fields, the length and direction of the crura, and the orientation of the cardinalia, which are hitherto undescribed. Three-dimensional reconstructions of articulated shells of N. pygmaea occurring in a single horizon were produced to show the orientation and length of arcuiform crura. The preservation of internal moulds together with the three-dimensional reconstruction of the internal shell morphology allow a more precise description of the internal morphology of this taxon than it is possible with articulated shells and serial sections. doi:10.1002/mmng.201200005

  13. Multigene analysis of lophophorate and chaetognath phylogenetic relationships. (United States)

    Helmkampf, Martin; Bruchhaus, Iris; Hausdorf, Bernhard


    Maximum likelihood and Bayesian inference analyses of seven concatenated fragments of nuclear-encoded housekeeping genes indicate that Lophotrochozoa is monophyletic, i.e., the lophophorate groups Bryozoa, Brachiopoda and Phoronida are more closely related to molluscs and annelids than to Deuterostomia or Ecdysozoa. Lophophorates themselves, however, form a polyphyletic assemblage. The hypotheses that they are monophyletic and more closely allied to Deuterostomia than to Protostomia can be ruled out with both the approximately unbiased test and the expected likelihood weights test. The existence of Phoronozoa, a putative clade including Brachiopoda and Phoronida, has also been rejected. According to our analyses, phoronids instead share a more recent common ancestor with bryozoans than with brachiopods. Platyhelminthes is the sister group of Lophotrochozoa. Together these two constitute Spiralia. Although Chaetognatha appears as the sister group of Priapulida within Ecdysozoa in our analyses, alternative hypothesis concerning chaetognath relationships could not be rejected.

  14. Impact Theory of Mass Extinctions and the Invertebrate Fossil Record (United States)

    Alvarez, Walter; Kauffman, Erle G.; Surlyk, Finn; Alvarez, Luis W.; Asaro, Frank; Michel, Helen V.


    There is much evidence that the Cretaceous-Tertiary boundary was marked by a massive meteorite impact. Theoretical consideration of the consequences of such an impact predicts sharp extinctions in many groups of animals precisely at the boundary. Paleontological data clearly show gradual declines in diversity over the last 1 to 10 million years in various invertebrate groups. Reexamination of data from careful studies of the best sections shows that, in addition to undergoing the decline, four groups (ammonites, cheilostomate bryozoans, brachiopods, and bivalves) were affected by sudden truncations precisely at the iridium anomaly that marks the boundary. The paleontological record thus bears witness to terminal-Cretaceous extinctions on two time scales: a slow decline unrelated to the impact and a sharp truncation synchronous with and probably caused by the impact.

  15. The post-Laramide clastic deposits of the Sierra de Guanajuato: Compositional implications on the tectono-sedimentary and paleographic evolution

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    Miranda-Aviles, R.; Puy-Alquiza, M.J.; OmaNa, L.; Loza-Aguirre, I.


    This article presents the results of the study on sedimentation, sedimentary environments, tectono-sedimentary and paleogeographic evolution of post-Laramide clastic deposits and pre-volcanism of the Sierra Madre Occidental in the Sierra de Guanajuato, central Mexico. The Eocene Duarte Conglomerate and Guanajuato Conglomerate were deposited in the middle and distal parts of alluvial fans. The studied rocks are composed of limestone clasts, granite, andesite, metasediments, diorite, and pyroxenite, indicating the erosion of uplifted blocks of the basal complex of the Sierra de Guanajuato (Arperos basin). The petrographic and compositional analysis of limestone shows a textural variation from basin limestones and shallow platform limestones. The shallow platform limestone contain bivalves, brachiopods, gastropods, echinoderms and benthic foraminifera from the Berriasian-Valanginian. The shallow-water limestone corresponds to the boundary of the Arperos basin whose original outcrops currently not outcrop in the Sierra de Guanajuato. (Author)

  16. A phylogenomic profile of hemerythrins, the nonheme diiron binding respiratory proteins

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    Mizuguchi Kenji


    Full Text Available Abstract Background Hemerythrins, are the non-heme, diiron binding respiratory proteins of brachiopods, priapulids and sipunculans; they are also found in annelids and bacteria, where their functions have not been fully elucidated. Results A search for putative Hrs in the genomes of 43 archaea, 444 bacteria and 135 eukaryotes, revealed their presence in 3 archaea, 118 bacteria, several fungi, one apicomplexan, a heterolobosan, a cnidarian and several annelids. About a fourth of the Hr sequences were identified as N- or C-terminal domains of chimeric, chemotactic gene regulators. The function of the remaining single domain bacterial Hrs remains to be determined. In addition to oxygen transport, the possible functions in annelids have been proposed to include cadmium-binding, antibacterial action and immunoprotection. A Bayesian phylogenetic tree revealed a split into two clades, one encompassing archaea, bacteria and fungi, and the other comprising the remaining eukaryotes. The annelid and sipunculan Hrs share the same intron-exon structure, different from that of the cnidarian Hr. Conclusion The phylogenomic profile of Hrs demonstrated a limited occurrence in bacteria and archaea and a marked absence in the vast majority of multicellular organisms. Among the metazoa, Hrs have survived in a cnidarian and in a few protostome groups; hence, it appears that in metazoans the Hr gene was lost in deuterostome ancestor(s after the radiata/bilateria split. Signal peptide sequences in several Hirudinea Hrs suggest for the first time, the possibility of extracellular localization. Since the α-helical bundle is likely to have been among the earliest protein folds, Hrs represent an ancient family of iron-binding proteins, whose primary function in bacteria may have been that of an oxygen sensor, enabling aerophilic or aerophobic responses. Although Hrs evolved to function as O2 transporters in brachiopods, priapulids and sipunculans, their function in

  17. Prolonged Permian Triassic ecological crisis recorded by molluscan dominance in Late Permian offshore assemblages. (United States)

    Clapham, Matthew E; Bottjer, David J


    The end-Permian mass extinction was the largest biotic crisis in the history of animal life, eliminating as many as 95% of all species and dramatically altering the ecological structure of marine communities. Although the causes of this pronounced ecosystem shift have been widely debated, the broad consensus based on inferences from global taxonomic diversity patterns suggests that the shift from abundant brachiopods to dominant molluscs was abrupt and largely driven by the catastrophic effects of the end-Permian mass extinction. Here we analyze relative abundance counts of >33,000 fossil individuals from 24 silicified Middle and Late Permian paleocommunities, documenting a substantial ecological shift to numerical dominance by molluscs in the Late Permian, before the major taxonomic shift at the end-Permian mass extinction. This ecological change was coincident with the development of fluctuating anoxic conditions in deep marine basins, suggesting that numerical dominance by more tolerant molluscs may have been driven by variably stressful environmental conditions. Recognition of substantial ecological deterioration in the Late Permian also implies that the end-Permian extinction was the climax of a protracted environmental crisis. Although the Late Permian shift to molluscan dominance was a pronounced ecological change, quantitative counts of 847 Carboniferous-Cretaceous collections from the Paleobiology Database indicate that it was only the first stage in a stepwise transition that culminated with the final shift to molluscan dominance in the Late Jurassic. Therefore, the ecological transition from brachiopods to bivalves was more protracted and complex than their simple Permian-Triassic switch in diversity.

  18. Prolonged Permian–Triassic ecological crisis recorded by molluscan dominance in Late Permian offshore assemblages (United States)

    Clapham, Matthew E.; Bottjer, David J.


    The end-Permian mass extinction was the largest biotic crisis in the history of animal life, eliminating as many as 95% of all species and dramatically altering the ecological structure of marine communities. Although the causes of this pronounced ecosystem shift have been widely debated, the broad consensus based on inferences from global taxonomic diversity patterns suggests that the shift from abundant brachiopods to dominant molluscs was abrupt and largely driven by the catastrophic effects of the end-Permian mass extinction. Here we analyze relative abundance counts of >33,000 fossil individuals from 24 silicified Middle and Late Permian paleocommunities, documenting a substantial ecological shift to numerical dominance by molluscs in the Late Permian, before the major taxonomic shift at the end-Permian mass extinction. This ecological change was coincident with the development of fluctuating anoxic conditions in deep marine basins, suggesting that numerical dominance by more tolerant molluscs may have been driven by variably stressful environmental conditions. Recognition of substantial ecological deterioration in the Late Permian also implies that the end-Permian extinction was the climax of a protracted environmental crisis. Although the Late Permian shift to molluscan dominance was a pronounced ecological change, quantitative counts of 847 Carboniferous–Cretaceous collections from the Paleobiology Database indicate that it was only the first stage in a stepwise transition that culminated with the final shift to molluscan dominance in the Late Jurassic. Therefore, the ecological transition from brachiopods to bivalves was more protracted and complex than their simple Permian–Triassic switch in diversity. PMID:17664426

  19. Hirnantian (latest Ordovician bio- and chemostratigraphy of the Stirnas-18 core, western Latvia

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    Hints, Linda


    Full Text Available Integrated study of the uppermost Ordovician Porkuni Stage in the Stirnas-18 core, western Latvia, has revealed one of the most complete Hirnantian successions in the eastern Baltic region. The interval is characterized by two shallowing upwards depositional sequences that correspond to the Kuldiga and Saldus formations. The whole-rock carbon stable isotope curve indicates a long rising segment of the Hirnantian carbon isotope excursion, with the highest peak in the upper part of the Kuldiga Formation. The bioclast carbon and oxygen curves fit well with the whole-rock carbon data. Micro- and macrofossil data enabled seven combined associations to be distinguished within the Hirnantian strata. The early Porkuni fauna of the Spinachitina taugourdeaui Biozone, with pre-Hirnantian affinities, is succeeded by an interval with a Hindella–Cliftonia brachiopod association, a specific polychaete fauna, the chitinozoan Conochitina scabra, and the conodont Noixodontus girardeauensis. The middle part of the Kuldiga Formation is characterized by a low-diversity Dalmanella testudinaria brachiopod association, high diversity of scolecodonts, and the occurrence of the chitinozoan Lagenochitina prussica. From the middle part of the Kuldiga Formation the youngest occurrence yet known of the conodont Amorphognathus ordovicicus is reported. Also typical of the Kuldiga Formation is the occurrence of the trilobite Mucronaspis mucronata. The uppermost Hirnantian Saldus Formation contains no shelly fauna, but yields redeposited conodonts and at least partly indigenous chitinozoans and scolecodonts. Palaeontological criteria and stable isotope data enable correlation of the Stirnas section with other Hirnantian successions in the Baltic region and elsewhere.

  20. Nineteenth-century collapse of a benthic marine ecosystem on the open continental shelf. (United States)

    Tomašových, Adam; Kidwell, Susan M


    The soft-sediment seafloor of the open continental shelf is among the least-known biomes on Earth, despite its high diversity and importance to fisheries and biogeochemical cycling. Abundant dead shells of epifaunal suspension-feeding terebratulid brachiopods ( Laqueus ) and scallops on the now-muddy mainland continental shelf of southern California reveal the recent, previously unsuspected extirpation of an extensive offshore shell-gravel ecosystem, evidently driven by anthropogenic siltation. Living populations of attached epifauna, which formerly existed in a middle- and outer-shelf mosaic with patches of trophically diverse muds, are restricted today to rocky seafloor along the shelf edge and to the sandier shelves of offshore islands. Geological age-dating of 190 dead brachiopod shells shows that (i) no shells have been produced on the mainland shelf within the last 100 years, (ii) their shell production declined steeply during the nineteenth century, and (iii) they had formerly been present continuously for at least 4 kyr. This loss, sufficiently rapid (less than or equal to 100 years) and thorough to represent an ecosystem collapse, coincides with intensification of alluvial-plain land use in the nineteenth century, particularly livestock grazing. Extirpation was complete by the start of twentieth-century urbanization, warming, bottom fishing and scientific surveys. The loss of this filter-feeding fauna and the new spatial homogeneity and dominance of deposit- and detritus-feeders would have altered ecosystem functioning by reducing habitat heterogeneity and seawater filtering. This discovery, attesting to the power of this geological approach to recent ecological transitions, also strongly increases the spatial scope attributable to the negative effects of siltation, and suggests that it has been under-recognized on continental shelves elsewhere as a legacy of coastal land use. © 2017 The Author(s).

  1. La moneda como ofrenda en los manantiales

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    Manuel Abad Varela


    aguas termales de Albula ^'. De Nerón nos dice que cuando reconstruyó su casa en Roma después del incendio, la famosa domus áurea, hizo llegar a las salas de baño agua de mar y de Albula *"*. Por las aras recogidas en las fuentes termales, sabemos que las divinidades que más se sintieron favorecidas con estas ofrendas monetales fueron las Ninfas y las aguas que más beneficios causaron o por las que se sintieron más agradecidos los visitantes fueron las de aguas sulfuradas- cálcicas, es decir, las que se recomiendan principalmente para los problemas de dermatosis herpética, neurosis y catarros crónicos de las vías respiratorias ^^ sin que ésto signifique que hubiese alguna relación entre las cualidades de las aguas y las divinidades a quienes se dedicaron las aras. Finalmente, conviene señalar que son éstas las únicas conclusiones a las que nos atrevemos a llegar partiendo de las informaciones que tenemos. No obstante, deseamos que en un futuro se produzcan más hallazgos en lugares tan particulares como los señalados, fuentes, ríos y lagos, que nos permitan confirmar o desmentir con más precisión nuestras hipótesis. Para que ésto suceda animo desde aquí a los arqueólogos para que busquen este tipo de yacimientos y tengan en cuenta sus ofrendas, tratándolas con cuidado por su mal estado, y no tardando en darlas a conocer como tales.This paper deals with the finding of thirty one cases of possible monetary offerings to the divinities in the waters of the Spanish península. The largest number of cases involve springs, which make up 74.19 % of the total, of which 78.26 % are hyperthermal springs with temperatures ranging between 15 and 70 C. Most of these springs are to be found ín the West of Spaín. They are mainly connected with the Nymphis, except in the North West, where they are offered up to Apollini, to judge from the devotional alters which can be sean. It would appear, from the coins collected, that the custom of throwing stipes to the

  2. Macroinvertebrate palaeo-communities from the Jurassic succession of Gebel Maghara (Sinai, Egypt) (United States)

    Abdelhady, Ahmed Awad; Fürsich, Franz Theodor


    Macrobenthic palaeo-communities of the Middle and Upper Jurassic strata of G. Maghara, Egypt, were investigated to identify relationships with environmental parameters and to trace the temporal changes of the ecosystem associated with sea-level fluctuations. The quantitative analysis of a data matrix comprising 198 macrobenthic taxa in 138 samples collected from four sections identified nine associations and three assemblages, interpreted to be representative of their original environment. Non-Metric Multidimensional Scaling (NMDS) delineated the same degree of habitat partitioning as hierarchical clusters with very little overlap. Detrended Correspondence Analysis (DCA) identified water depth as the primary environmental gradient controlling the distribution of the fauna, while Axis 2 reflects substrate consistency. Community structure is related to the various ramp environments. Based on diversities, the associations and assemblages have been divided into two major groups, low-stress polyspecific associations and high-stress paucispecific associations. The low-stress polyspecific associations were interpreted to represent two different habitats, a high-energy, firm substrate habitat, in which epifaunal bivalves and brachiopods in addition to solitary corals dominated during advanced stages of transgression, and a low-energy, soft substrate habitat dominated by infaunal bivalves during the maximum flooding. The high-stress paucispecific associations are dominated by one or few taxa and occurred (1) in an oligotrophic setting that developed during episodes of sediment starvation in restricted inner ramp environments or during early transgression, (2) in a setting characterized by high sedimentation rates which developed during advanced regression, (3) in a distal prodelta setting with soft substrate and dysoxia during sea-level lowstand, and (4) in a high-energy shoal environment during peak regression. A combined stress involving a shortage in food supply

  3. A review of the Late Cambrian (Furongian) palaeogeography in the western Mediterranean region, NW Gondwana (United States)

    Álvaro, J. Javier; Ferretti, Annalisa; González-Gómez, Cristina; Serpagli, Enrico; Tortello, M. Franco; Vecoli, Marco; Vizcaïno, Daniel


    , onchonotinids, and pagodiids), linguliformean brachiopods (acrotretids, obolids, scaphelasmatids, siphonotretids, and zhanatellids), echinoderms (mitrates, glyptocystitid cystoids, and stromatocystoids), and conodonts belonging to the lower Peltura Zone; and (iii) the subsequent input of new trilobites (asaphids, calymenids, catillicephalids, nileids and remopleurids), which marks the base of the Proteuloma geinitzi Zone, associated with pelmatozoan holdfasts ( Oryctoconus), and a distinct input of late Tremadocian conodonts ( Paltodus deltifer Zone). The biogeographic distribution of latest Middle and Late Cambrian trilobites supports brachiopod data indicating strong affinities between the western Mediterranean region, East Gondwana (North China/Korea, South China, Australia, and Antarctica) and Kazakhstania during the late Languedocian, which became significantly stronger during the Late Cambrian. This major shift may suggest modification in oceanic circulation patterns throughout Gondwana across the Middle-Late Cambrian transition.

  4. Episodic events in long-term geological processes: A new classification and its applications

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    Dmitry A. Ruban


    Full Text Available Long-term geological processes are usually described with curves reflecting continuous changes in the characteristic parameters through the geological history, and such curves can be employed directly for recognition of episodic (relatively long-term events linked to these changes. The episodic events can be classified into several categories according to their scale (ordinary and anomalous events, “shape” (positive, negative, and neutral events, and relation to long-term trend change (successive, interruptive, facilitative, stabilizing, transformative, increasing, and decreasing. Many types of these events can be defined depending on the combination of the above-mentioned patterns. Of course, spatial rank, duration, and origin can be also considered in description of these events. The proposed classification can be applied to events in some real long-term geological processes, which include global sea-level changes, biodiversity dynamics, lithospheric plate number changes, and palaeoclimate changes. Several case examples prove the usefulness of the classification. It is established that the Early Valanginian (Early Cretaceous eustatic lowstand (the lowest position of the sea level in the entire Cretaceous was negative, but ordinary and only interruptive event. In the other case, it becomes clear that the only end-Ordovician and the Permian/Triassic mass extinctions transformed the trends of the biodiversity dynamics (from increase to decrease and from decrease to increase respectively, and the only Cretaceous/Paleogene mass extinction was really anomalous event on the Phanerozoic biodiversity curve. The new palaeontological data are employed to reconstruct the diversity dynamics of brachiopods in Germany (without the Alps and the Swiss Jura Mountains. The further interpretation of the both diversity curves implies that the Early Toarcian mass extinction affected the regional brachiopod faunas strongly, but this event was only decreasing

  5. Patterns in bioclastic accumulation through the Phanerozoic: Changes in input or in destruction? (United States)

    Kidwell, Susan M.; Brenchley, Patrick J.


    Evolutionary changes in the ecology and diversity of organisms that produce and destroy calcareous skeletons suggest that bioclastic concentrations themselves might have changed in nature through the Phanerozoic. Empirical data from marine siliciclastic records of Ordovician-Silurian, Jurassic, and Neogene ages indicate a significant increase in the thickness of densely packed bioclastic concentrations over geologic time, from a primarily thin-bedded brachiopod-dominated record in the Ordovician-Silurian to a mollusk-dominated record with many more and thicker shell beds in the Neogene. Jurassic shell beds vary in thickness with the Paleozoic or modern affinities of the chief constituents, suggesting, along with other evidence, that the Phanerozoic increase was determined neither by diagenesis nor by a shift in taphonomic conditions on the sea floor but rather by the evolution of bioclast producers, namely, groups with (1) more durable low-organic skeletons, (2) greater ecological success in high-energy habitats, and (3) on the basis of indirect evidence, higher rates of carbonate production. These results suggest that (1) reproductive and metabolic output has increased in benthic communities over time and (2) the scale of time averaging in benthic assemblages has increased owing to greater hard-part durability of modern groups.

  6. Anoxia, toxic metals and acidification: volcanically-driven causes of the Middle Permian (Capitanian) mass extinction in NW Pangaea? (United States)

    Bond, David; Grasby, Stephen; Wignall, Paul


    The controversial Capitanian (Middle Permian, 262 Ma) mass extinction, mostly known from equatorial latitudes, has recently been identified in a Boreal setting in Spitsbergen. We now document this extinction in the record of brachiopods from the Sverdrup Basin in NW Pangaea (Ellesmere Island, Canada), confirming Middle Permian losses as a global crisis on par with the "Big Five". Redox proxies (pyrite framboids and trace metals) show that the high latitude crisis coincided with an intensification of oxygen-poor conditions - a potent killer that is not clearly developed in lower latitude sections. Mercury becomes briefly enriched in strata at the level of the Middle Permian extinction level in Spitsbergen and Ellesmere Island, indicating voluminous but short-lived volcanism that is likely to have been the emplacement of the Emeishan large igneous province (LIP) in SW China. A potent cocktail of poisons appears to have impacted across the Boreal Realm, whilst the near-total loss of carbonates near the extinction level is also consistent with reduced pH across the region. Multiple stresses, possibly with origins in low-latitude LIP volcanism, are therefore implicated in the Middle Permian extinction and there was no respite even in the far-distant Boreal Realm.

  7. Benthic biodiversity and ecological gradients in the Seno Magdalena (Puyuhuapi Fjord, Chile) (United States)

    Betti, F.; Bavestrello, G.; Bo, M.; Enrichetti, F.; Loi, A.; Wanderlingh, A.; Pérez-Santos, I.; Daneri, G.


    Due to its complex hydrological, geomorphological and climatic features, the Chilean fjords region is considered among the most productive areas of the world. The benthic fauna of this region accounts for more than 1600 species showing marked latitudinal biogeographic differences characterizing this as one of the most important hotspot of biodiversity of cold-temperate environments. Despite numerous studies have been conducted to depict the biological characteristics of the fjords, the present situation is strongly unbalanced towards specific taxa. Hence, this study takes into consideration a community approach, highlighting the distribution of six benthic assemblages thriving on vertical walls along the Seno Magdalena fjord (Aysen region). Underwater pictures were used to characterize the trends in abundance and diversity of the main taxa showing distinct responses to salinity and turbidity. Among the less tolerant taxa to high fresh water inputs there are encrusting algae, mainly found in the most external sites lashed by outer currents, far from the estuarine plume. The bathymetric zonation of the assemblages, instead, is characterized by a dense mussel belt in the first 10 m, within a thick layer of low-salinity, nutrient-enriched waters. Rich assemblages of sponges, brachiopods, gorgonians and scleractinians thrive in deeper, marine, clear waters. The evaluation of the ecological role of benthic species leads both to the definition of potential bioindicator taxa responding to anthropic disturbances and to the promotion of protected areas.


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    Full Text Available The Arroyo Malo Formation at Alumbre Creek, on the northern bank of the Atuel River, west central Argentina, comprises a c. 300 m thick continuous marine succession across the Triassic-Jurassic System boundary, consisting of massive and laminated pelites indicative of a slope depositional environment. Late Triassic invertebrates, including ammonoids, nautiloids, bivalves, gastropods, brachiopods and corals are restricted to the lower 150 m. Beds between 125-135 m from the bottom yield Choristoceras cf. marshi Hauer, a species found in the Marshi/Crickmayi Zone of Europe and North America, together with loose fragments of Psiloceras cf. pressum Hillebrandt, coeval with the lower to middle part of the Hettangian Planorbis Zone. About 80 m higher are beds yielding Psiloceras cf. rectocostatum Hillebrandt, a species that gives name to an Andean biozone partially coeval with the Johnstoni and Plicatulum Subzones, upper Planorbis Zone. Other fossils recorded in the Rhaetian strata of this section are foraminifers, ostracods and plant remains identified as Zuberia cf. zuberi (Szaj. Freng. and Clathropteris sp. The section was also sampled for conodonts and radiolarians, thus far with negative results. A palaeomagnetic study is underway.

  9. The fossil record, function, and possible origins of shell color patterns in Paleozoic marine invertebrates

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    Kobluk, D.R. (Univ. of Toronto, Ontario (Canada)); Mapes, R.H. (Ohio Univ., Athens (USA))


    Fossil invertebrate shells and carapaces displaying preserved original color patterns are among the rarest fossils. The fossil record of color patterns extends into the Middle Cambrian where the trilobite Anomocare displays a fan-like array of stripes on the pygidium. About 180 Paleozic genera are known with patterns, including trilobites, cephalopods, gastropods, brachiopods, bivalves, crinoids, and crustaceans. Based upon an analysis of these taxa, it appears that patterns and pigments in middle and late Paleozoic invertebrates may have served several functions such as warning displays, light screening, camouflage, or waste disposal. However, the presence of color patterns in fossil invertebrates in the early Paleozoic may have developed prior to the evolution of vision sufficiently sophisticated to see them. This suggests that camouflage and warning displays were not the original functions of color patterns, and that in the earliest Paleozoic they may not have been functional. The authors propose a hypothesis that involves three developmental phases in the evolution of invertebrate color patterns: (1) the incorporation of metabolic by-products, perhaps some pigmented and some not pigmented, into shells and carapaces as a means of disposal of dietary or metabolic wastes, (2) use of these pigments and patterns as an environmental adaptation, such as light screening, and (3) display during and following the evolution of vision in predators sufficiently sophisticated to see the patterns.

  10. Comparison of two carbonate mound sequences in the Lower Ordovician El Paso Formation, west Texas and southern New Mexico

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    Clemons, R.E.


    The El Paso Formations consists of four members, in ascending order: Hitt Canyon, Jose McKelligon and Padre. Mounds in the McKelligon Member exposed in the southern Franklin Mountains were described by Toomey (1970). Most of these mounds are small but one large one is 5.8 m thick and about 13.7 m long in outcrop. The mound rock is chiefly bioclastic wackestone with minor packstone and boundstone. The varied fauna contains echinoderms, sponges and spicules, gastropods, trilobites, digitate algae, Nuia, Girvanella, Pulchrilamina, Calathium, and minor brachiopods and cephalopods. Intraclastic, bioclastic grainstone fills channels cut in the mounds. Similar, but smaller and less spectacular mounds occur in the McKelligon Member in the Florida, Big Hatchet, and Caballo Mountains, Lone Mountain, Cooke's Range, and elsewhere in southwestern New Mexico. A second type of mound is common in the upper part of the Hitt Canyon Member in the Cooke's Range, Red Hills, Caballo and Big Hatchet Mountains. These mounds also are typically small but one in the Red Hills is 13.7 m thick and about 30 m long in outcrop. The mound complex is about 75-80% SH-C and LLH-C stromatolite boundstone and bioclastic wackestone. The remaining 20-25% is bioclastic packstone and grainstone between the SH-C stromatolites and filling channels cut in the mound complex. The limited fauna contains small fragments of echinoderms, gastropods, trilobites, spicules, and Nuia.

  11. Late Ordovician pelecypod faunas from the Cincinnati, Ohio area

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    Frey, R.C.


    The distribution of pelecypod faunas in the Late Ordovician strata exposed in the Cincinnati, Ohio area, points to a close relationship between lithofacies type and the life habits of these Ordovician bivalves. Muddy clastic shallow marine facies of Edenian, Maysvillian, and early Richmondian age support faunas dominated by endobyssate filter-feeding species, including a variety of modiomorphids and the genus Ambonychia, plus infaunal filter-feeding orthonotids, and in faunal deposit-feeding palaeotaxodonts. These pelecypod groups occur in claystones with a fauna of calymenid and asaphid trilobites, nautiloids, cyclomyan monoplacophorans, and occasionally crinoids and asterozoans. Younger Richmondian strata in the area are predominantly carbonate platform facies and support pelecypod faunas dominated by robust endobyssate and epibyssate ambonychiids, cyrtodontids, and colpomyids. These pelecypods are associated with diverse assemblage of articulate brachiopods, trepostome ectoprocts, solitary rugose corals, and mollusks in skeletal limestones representing storm-reworked thickets or ramos ectoprocts. This fundamental dichotomy in Late Ordovician pelecypod faunas is recognized not only in the Cincinnati area, but in Late Ordovician strata exposed on Manitoulin Island in Ontario and eastward into Quebec. Reconstructions of the life habits of these pelecypods demonstrates the dominance of the endobyssate mode of life in these Early Paleozoic pelecypods.

  12. Global iridium anomaly, mass extinction, and redox change at the Devonian-Carboniferous boundary

    Energy Technology Data Exchange (ETDEWEB)

    Wang, K. (Geological Survey of Canada, Calgary, Alberta (Canada) Univ. of Calgary, Alberta (Canada)); Attrep, M. Jr.; Orth, C.J. (Los Alamos National Lab., NM (United States))


    Iridium abundance anomalies have been found on a global scale in the Devonian-Carboniferous (D-C) boundary interval, which records one of the largest Phanerozoic mass-extinction events, an event that devastated many groups of living organisms, such as plants, ammonoids, trilobites, conodonts, fish, foraminiferans, brachiopods, and ostracodes. At or very close to the D-C boundary, there exists a geographically widespread black-shale interval, and Ir abundances reach anomalous maxima of 0.148 ppb (Montagne Noire, France), 0.138 ppb (Alberta, Canada) 0.140 ppb (Carnic Alps, Austria), 0.156 ppb (Guangxi, China), 0.258 ppb (Guizhou, China), and 0.250 ppb (Oklahoma). The discovery of global D-C Ir anomalies argues for an impact-extinction model. However, nonchondritic ratios of Ir to other important elements and a lack of physical evidence (shocked quartz, microtektites) do not support such a scenario. The fact that all Ir abundance maxima are at sharp redox boundaries in these sections leads us to conclude that the Ir anomalies likely resulted from a sudden change in paleo-redox conditions during deposition and/or early diagenesis. 36 refs., 2 figs., 1 tab.

  13. Depositional model for Rival and Midale subintervals (Mississippian), north-central Burke County, North Dakota

    Energy Technology Data Exchange (ETDEWEB)

    Harris, T.L.


    The Rival and Midale subintervals (Charles Formation, Upper Mississippian), north-central Burke County, North Dakota, represent two relative sea level fluctuations. Updip (northeast), the Rival subinterval contains fine to medium-bedded and chicken-wire anhydrite with interbedded algal bindstone that was deposited on supratidal flats. Basinward (southwest), the lithology changes to oncolitic, peloidal, intraclastic grainstone/packstone that was deposited in intertidal and subtidal restricted lagoonal environments. Evaporites precipitated in the sediment of the intertidal to shallow subtidal restricted lagoonal environment. Overlying the Rival subinterval is skeletal wackestone and packstone of the lower Midale subinterval. The presence of normal-marine fauna (crinoids, brachiopods, trilobites, rugose and tabulate coral) indicates a significant relative sea level transgression occurred following deposition of the Rival. The middle and upper Midale subinterval consists of intensely burrowed dolowackestone and dolomudstone that contain a less diversified faunal assemblage. Overlying the Midale carbonates is a transitional zone of calcareous shale and dolomite that grades upward into mottled (burrowed.) and finely laminated microgranular dolomite and anhydrite. The upper Midale section represents a relative sea level regression (shoreline progradation). Updip (northeast) reservoirs produce from the Midale carbonates, which are sealed laterally and vertically by calcarous shale and microgranular dolomitic anhydrite of the Midale Evaporite. Downdip (southwest), the Rival produces from porous grainstone, which is sealed laterally by intertidal/supratidal carbonates and evaporites, resulting in a stratigraphic trap. Vuggy and intergranular porosity are the major porosity types in the Rival grainstone, and moldic and intercrystalline porosity are dominant in the Midale dolowackestone.

  14. Measuring the eyes of trilobites: a motorized goniometer for macropaleontology

    Energy Technology Data Exchange (ETDEWEB)

    Cowen, R.; Cooper, J.H.


    The geometry of the trilobite visual surface is important in studies of trilobite taxonomy, paleoecology, and evolution. Unfortunately, methods for measuring visual surface features have been slow and laborious. The authors have built an apparatus which allows rapid, reliable measurement of angular relationships on the trilobite cephalon. A specimen is mounted on a set of turntables rotated by steeper motors driven by impulses from a computer terminal. As the turntables are rotated, their angular displacements from zero are automatically recorded to a repeatable accuracy of less than half a degree. Thus the angular positions of lens axes, spines, glabella, and other important features can be recorded directly into the computer for further processing and graphics display. The authors apparatus was designed to study the eyes of phacopid-trilobites, for two reasons. First, the Phacops rana group of trilobites is one of the type examples of punctuated speciation. Second, it has been suggested that phacopids had stereoscopic vision in each eye. The new apparatus will allow rapid and accurate study of suites of specimens, in order to test these hypotheses. Their new apparatus, essentially a motorized goniometer, could be used or modified to work with any specimens in which rapid goniometric measurements are useful. Such applications could include routine morphometric studies of molluscan or brachiopod shells or echinoderm tests, as well as specific projects dealing with spines, pores, plates, or septa in a wide variety of fossil invertebrates.

  15. Statistically significant faunal differences among Middle Ordovician age, Chickamauga Group bryozoan bioherms, central Alabama

    Energy Technology Data Exchange (ETDEWEB)

    Crow, C.J.


    Middle Ordovician age Chickamauga Group carbonates crop out along the Birmingham and Murphrees Valley anticlines in central Alabama. The macrofossil contents on exposed surfaces of seven bioherms have been counted to determine their various paleontologic characteristics. Twelve groups of organisms are present in these bioherms. Dominant organisms include bryozoans, algae, brachiopods, sponges, pelmatozoans, stromatoporoids and corals. Minor accessory fauna include predators, scavengers and grazers such as gastropods, ostracods, trilobites, cephalopods and pelecypods. Vertical and horizontal niche zonation has been detected for some of the bioherm dwelling fauna. No one bioherm of those studied exhibits all 12 groups of organisms; rather, individual bioherms display various subsets of the total diversity. Statistical treatment (G-test) of the diversity data indicates a lack of statistical homogeneity of the bioherms, both within and between localities. Between-locality population heterogeneity can be ascribed to differences in biologic responses to such gross environmental factors as water depth and clarity, and energy levels. At any one locality, gross aspects of the paleoenvironments are assumed to have been more uniform. Significant differences among bioherms at any one locality may have resulted from patchy distribution of species populations, differential preservation and other factors.

  16. Possible extrinsic controls on the Ordovician radiation: Stratigraphic evidence from the Great Basin, western USA

    Energy Technology Data Exchange (ETDEWEB)

    Droser, M.L. (Univ. of California, Riverside, CA (United States). Dept. of Earth Sciences); Fortey, R.A. (Natural History Museum, London (United Kingdom). Dept. of Palaeontology)


    The Ordovician radiation has been previously examined by looking at 1/analyses of patterns of diversification within small clades, 2/analyses of large databases to elucidate large-scale paleoecological patterns such as increased tiering and onshore-offshore shifts associated with this radiation. In order to resolve the relationships between these two scales of analysis there is critical need to examine in detail the paleoecology and possible biofacies shifts associated with the Ordovician radiation. The authors have examined the base of the Whiterock Series (Lower-Middle Ordovician) in the Great Basin as it represents one of the most complete records of the Ordovician radiation on the North American continent. Detailed field evidence suggests that the base of the Whiterock does not represent a simple faunal turnover but corresponds with the first occurrences in the region of groups that come to dominate the rest of the Paleozoic. Among the trilobites, this includes the lichides, calymenids, proetides, and phacopides. Similar patterns are found among the dominate Paleozoic bivalve, cephalopod, brachiopod and graptolite clades. Global correlation of this time interval suggests that this pattern of first broad geographic occurrences is not unique to North America. This boundary corresponds with a globally recognized sea level lowstand. In the Great Basin, significant facies shifts are present in shallow and deep water settings. While extrinsic controls are commonly reserved for extinctions, these data suggest that extrinsic factors may have been significant in the timing of the Paleozoic fauna rose to dominance.

  17. Les séries du carbonifère inférieur de la région d'Adarouch, NE du Maroc central: lithologie et biostratigraphieEarly carboniferous series of the Adarouch area, northeast central Morocco: lithology and biostratigraphy (United States)

    Berkhli, M.; Vachard, D.; Paicheler, J.-C.


    The early Carboniferous series of the Adarouch area (northeast central Morocco) are subdivided into three sedimentological and biostratigraphical units. The first unit, which belongs to the Late Visean zones V3bβ and V3bγ, was deposited on shallow carbonate platforms. The second unit belongs to the Late Visean zone, V3c, and incudes terrigenous deposits, such as turbidites, shales and olistostromes. The third unit belongs to the Serpukhovian stage and consists of sandstones and limestones. A new biostratigraphical analysis, which is based on foraminiferal, algae and pseudoalgae, allows an accurate dating of the units. The deposits of the zones V3bβ and V3bγ contain characteristic calcareous microfossils, such as Stacheoides sp., Pseudoendothyra sp. and Ungdarella uralica. The V3c zone (300-400 m) is shown in two oolitic beds of the Mouarhaz and Akerchi Formations, respectively, with Janischewskina sp. and Asteroarchaediscus sp. The Serpukhovian stage is characterised by the disappearance of the algae Koninckopora and the appearance of the brachiopod Titanaria. The new data from the Adarouch area confirm the Moroccan biostratigraphical scale of the Moroccan meseta.

  18. Metabolic flexibility: the key to long-term evolutionary success in Bryozoa? (United States)

    Peck, Lloyd S; Barnes, David K A


    Oxygen consumption (MO2) and activity were evaluated in Antarctic Bryozoa. Three species representing two different morphologies, flat sheet, laminar forms, Isoseculiflustra tenuis and Kymella polaris, and the bush form Camptoplites bicornis were used. In Bryozoa, activity is measured as the proportion of colony zooids with their lophophores extended. In I. tenuis and K. polaris, residual analysis showed that the percentage of zooids with extended lophophores was not correlated with colony MO2. Lophophore extension is, therefore, a poor measure of activity, and other costs (e.g. growth, reproduction, storage) probably form the major metabolic costs. MO2 per unit of ash-free dry mass (AFDM) in the laminar forms was low compared with other Antarctic marine invertebrates, but not lower than brachiopods and echinoderms. However, the lowest rate here, 16.8 microg O2 g AFDM(-1) h(-1) for a K. polaris colony, is (to our knowledge) the lowest for any animal so far reported. MO2 per unit of AFDM for C. bicornis, however, is among the highest reported for sessile or slow moving Antarctic marine ectotherms, with values similar to those for bivalve and gastropod molluscs. The highest rate, 527 microg O2 g AFDM(-1) h(-1) for one colony is (to our knowledge) the highest reported for polar animals of this type. Extreme diversity in metabolic strategy may explain the bryozoan long evolutionary record and great success in shallow marine environments worldwide.

  19. The late Middle Devonian fauna of Red Hill I, Nevada, and its paleobiogeographic implications

    Directory of Open Access Journals (Sweden)

    H.-P. Schultze


    Full Text Available The fauna of the Middle Devonian Red Hill I locality, Nevada, is unusual in the co-occurrence of a rich fish assemblage with a rich invertebrate one. Sponges are second in abundance of specimens and number of species only to the fishes and occur together with other invertebrates (conodonts, conulariids, dacryoconarid tentaculites, gastropods, bivalves, brachiopods, arthropods, and unidentifiable ammonoids and echinoderms. The invertebrates indicate a marine depositional paleoenvironment. The conodonts indicate a placement within the lower disparalis Zone, late Givetian. The fish assemblage is dominated by the antiarch Asterolepis. All the other fishes, acanthodians, actinopterygians and sarcopterygians, are less common. The closest biogeographic relationship of the fish fauna is with the Middle/Late Devonian fish fauna of the Baltic Region, followed by that of eastern Canada (Miguasha, Scotland and Iran. This distribution corresponds to the Devonian Euramerica faunal province with connection to eastern Gondwana (Iran and Australia. Localities with the same genera as Red Hill I are interpreted as marine with the exception of the Scottish localities. Asterolepis is the most widely distributed vertebrate genus, mostly marine, but it may be able to enter freshwater like Eusthenopteron if one accepts a freshwater depositional paleoenvironment for the Scottish localities. doi:10.1002/mmng.201000001

  20. Structural changes of marine communities over the Permian-Triassic transition: Ecologically assessing the end-Permian mass extinction and its aftermath (United States)

    Chen, Zhong-Qiang; Tong, Jinnan; Liao, Zhuo-Ting; Chen, Jing


    The Permian/Triassic (P/Tr) transition is ecologically assessed based on examining 23 shelly communities from five shallow platform, ramp and shelf basin facies Permian-Triassic boundary (PTB) sections in South China. The shelly communities have undergone two major collapses coinciding with the two episodes of the end-Permian mass extinction. The first P/Tr extinction event devastated shelly communities in all types of settings to some extent. The basin communities have been more severely impacted than both platform and ramp communities. The survival faunas have rebounded more rapidly in shallow niches than in relatively deep habitats. The second P/Tr crisis destroyed the survival communities in shallow setting and had little impact on the basin communities in terms of community structures. The early Griesbachian communities are overall low-diversity and high-dominance. The governorship switch from brachiopods to bivalves in marine communities has been facilitated by two pulses of the end-Permian mass extinction and the whole takeover process took about 200 ka across the P/Tr boundary. Bivalve ecologic takeover initially occurred immediately after the first P/Tr extinction in shallow water habitats and was eventually completed in all niches after the second P/Tr event. Some post-extinction communities have the irregular rarefaction curves due to the unusual community structures rather than sampling intensities.

  1. Transcriptome analysis elucidates key developmental components of bryozoan lophophore development

    KAUST Repository

    Wong, Yue Him


    The most recent phylogenomic study suggested that Bryozoa (Ectoprocta), Brachiopoda, and Phoronida are monophyletic, implying that the lophophore of bryozoans, phoronids and brachiopods is a synapomorphy. Understanding the molecular mechanisms of the lophophore development of the Lophophorata clade can therefore provide us a new insight into the formation of the diverse morphological traits in metazoans. In the present study, we profiled the transcriptome of the Bryozoan (Ectoproct) Bugula neritina during the swimming larval stage (SW) and the early (4 h) and late (24 h) metamorphic stages using the Illumina HiSeq2000 platform. Various genes that function in development, the immune response and neurogenesis showed differential expression levels during metamorphosis. In situ hybridization of 23 genes that participate in the Wnt, BMP, Notch, and Hedgehog signaling pathways revealed their regulatory roles in the development of the lophophore and the ancestrula digestive tract. Our findings support the hypothesis that developmental precursors of the lophophore and the ancestrula digestive tract are pre-patterned by the differential expression of key developmental genes according to their fate. This study provides a foundation to better understand the developmental divergence and/or convergence among developmental precursors of the lophophore of bryozoans, branchiopods and phoronids.

  2. Lipid and fatty acid fractions in Lingula anatina (Brachiopoda: an intertidal benthic fauna in the West Bengal-Orissa coast, India

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    Samaresh Samanta


    Full Text Available Objective: To record the fractional components of lipid and polyunsaturated fatty acids of Lingula anatina (L. anatina, a Precambrian intertidal benthic brachiopod, giving emphasis on -ω series group especially eicosapentaenoic acid (EPA and docosahexaenoic acid (DHA alongside assessing their biotransformation within the population and mangrove-estuarine associated community. Methods: Different biological samples after being collected from three contrasting study sites viz. SI, SII and SIII at Talsari (Longitude 87°5′ E to 88°5′ E and Latitude 20°30′ N to 22°2′ N were stored at -20 °C until analyzed. Total lipids were extracted from each sample following Bligh and Dryer method. Identification and conformation of fatty acids were done by following Ackman method. Results: On analyzing different collected samples, muscles of L. anatina exhibited the highest amount of total lipids (2.95% of which 54.03% belongs to phospholipid groups. Different body parts of studied species contained appreciable and greater amount of EPA and DHA than α-linolenic acid. Conclusions: Different collected samples exhibited variabilities in respect of total lipids and its fractional fatty acid components. The muscles of L. anatina showed maximum storage of lipids and fatty acids. Differential occurrences of EPA and DHA in different body parts of L. anatina are supposed to be due to the biotransformation process converting the α-linolenic acid from its primary food sources.

  3. Proposed stratotype for the base of the highest Cambrian stage at the first appearance datum of Cordylodus andresi, Lawson Cove section, Utah, USA (United States)

    Miller, J.F.; Ethington, Raymond L.; Evans, K.R.; Holmer, L.E.; Loch, James D.; Popov, L.E.; Repetski, J.E.; Ripperdan, R.L.; Taylor, John F.


    We propose a candidate for the Global Standard Stratotype-section and Point (GSSP) for the base of the highest stage of the Furongian Series of the Cambrian System. The section is at Lawson Cove in the Ibex area of Millard County, Utah, USA. The marker horizon is the first appearance datum (FAD) of the conodont Cordylodus andresi Viira et Sergeyeva in Kaljo et al. [Kaljo, D., Borovko, N., Heinsalu, H., Khazanovich, K., Mens, K., Popov, L., Sergeyeva, S., Sobolevskaya, R., Viira, V., 1986. The Cambrian-Ordovician boundary in the Baltic-Ladoga clint area (North Estonia and Leningrad Region, USSR). Eesti NSV Teaduste Akadeemia Toimetised. Geologia 35, 97-108]. At this section and elsewhere this horizon also is the FAD of the trilobite Eurekia apopsis (Winston et Nicholls, 1967). This conodont characterizes the base of the Cordylodus proavus Zone, which has been recognized in many parts of the world. This trilobite characterizes the base of the Eurekia apopsis Zone, which has been recognized in many parts of North America. The proposed boundary is 46.7 m above the base of the Lava Dam Member of the Notch Peak Formation at the Lawson Cove section. Brachiopods, sequence stratigraphy, and carbon-isotope geochemistry are other tools that characterize this horizon and allow it to be recognized in other areas. ?? 2006 Nanjing Institute of Geology and Palaeontology, CAS.

  4. Megabenthic assemblages in the continental shelf edge and upper slope of the Menorca Channel, Western Mediterranean Sea (United States)

    Grinyó, Jordi; Gori, Andrea; Greenacre, Michael; Requena, Susana; Canepa, Antonio; Lo Iacono, Claudio; Ambroso, Stefano; Purroy, Ariadna; Gili, Josep-Maria


    Highly diverse megabenthic assemblages dominated by passive and active suspension feeders have been recently reported in shelf edge environments of the Mediterranean Sea. Due to their frequent association with species of commercial interest, these assemblages have been heavily impacted by fishing. The vulnerability and low resilience of these assemblages, composed mainly by long-living and slow-growing species, have motivated the implementation of management measures such as the restriction of bottom trawling, and the establishment of large protected areas embracing these environments. The Menorca Channel is one of such areas recently included in the European Union Natura 2000 network. Quantitative analysis of video transects recorded at 95-360 m depth by manned submersible and remotely operated vehicles were used to characterize megabenthic assemblages and to assess their geographical and bathymetric distribution. Six different assemblages were identified, mainly segregated by substrate type and depth. Hard substrates hosted coral gardens and sponge grounds, whereas soft sediments were mainly characterized by large extensions of the crinoid Leptometra phalangium and the brachiopod Gryphus vitreus. The good preservation of most of the observed assemblages is probably related to a low bottom trawling pressure, which mainly concentrates deeper on the adjacent continental slope. Because of their biological and ecological value, management and conservation measures need to be established to preserve these benthic assemblages.

  5. A review of phosphate mineral nucleation in biology and geobiology. (United States)

    Omelon, Sidney; Ariganello, Marianne; Bonucci, Ermanno; Grynpas, Marc; Nanci, Antonio


    Relationships between geological phosphorite deposition and biological apatite nucleation have often been overlooked. However, similarities in biological apatite and phosphorite mineralogy suggest that their chemical formation mechanisms may be similar. This review serves to draw parallels between two newly described phosphorite mineralization processes, and proposes a similar novel mechanism for biologically controlled apatite mineral nucleation. This mechanism integrates polyphosphate biochemistry with crystal nucleation theory. Recently, the roles of polyphosphates in the nucleation of marine phosphorites were discovered. Marine bacteria and diatoms have been shown to store and concentrate inorganic phosphate (Pi) as amorphous, polyphosphate granules. Subsequent release of these P reserves into the local marine environment as Pi results in biologically induced phosphorite nucleation. Pi storage and release through an intracellular polyphosphate intermediate may also occur in mineralizing oral bacteria. Polyphosphates may be associated with biologically controlled apatite nucleation within vertebrates and invertebrates. Historically, biological apatite nucleation has been attributed to either a biochemical increase in local Pi concentration or matrix-mediated apatite nucleation control. This review proposes a mechanism that integrates both theories. Intracellular and extracellular amorphous granules, rich in both calcium and phosphorus, have been observed in apatite-biomineralizing vertebrates, protists, and atremate brachiopods. These granules may represent stores of calcium-polyphosphate. Not unlike phosphorite nucleation by bacteria and diatoms, polyphosphate depolymerization to Pi would be controlled by phosphatase activity. Enzymatic polyphosphate depolymerization would increase apatite saturation to the level required for mineral nucleation, while matrix proteins would simultaneously control the progression of new biological apatite formation.

  6. Bioerosion and encrustation: Evidences from the Middle ‒ Upper Jurassic of central Saudi Arabia (United States)

    El-Hedeny, Magdy; El-Sabbagh, Ahmed; Al Farraj, Saleh


    The Middle ‒ Upper Jurassic hard substrates of central Saudi Arabia displayed considerable signs of bioerosion and encrustations. They include organic (oysters, other bivalves, gastropods, corals and brachiopods) and an inorganic carbonate hardground that marks the boundary between the Middle Jurassic Tuwaiq Mountain Limestone and the Upper Jurassic Hanifa Formation. Traces of bioerosion in organic substrates include seven ichnotaxa produced by bivalves (Gastrochaenolites Leymerie, 1842), polychaete annelids (Trypanites Mägdefrau, 1932; MaeandropolydoraVoigt, 1965 and CaulostrepsisClarke, 1908), sponges (Entobia Bronn, 1837), acrothoracican cirripedes (Rogerella Saint-Seine, 1951), gastropods (Oichnus Bromley, 1981) and probable ?Centrichnus cf. eccentricus. The encrusting epifauna on these substrates consist of several organisms, including oysters, serpulid worms, corals and foraminifera. In contrast, the carbonate hardground was only bioeroded by Gastrochaenolite, Trypanites and Entobia. Epibionts on this hardground include ;Liostrea Douvillé, 1904-type; oysters, Nanogyra nana Sowerby, 1822 and serpulids. In general, bioerosion and encrustation are less diversified in hardground than in organic substrates, indicating a long time of exposition of organic substrates with slow to moderate rate of deposition in a restricted marine environment. Both organic and inorganic commuinities are correlated with those of other equatorial, subtropical and temperate equivalents.

  7. Dolomitized bryozoan bioherms from the Lower Silurian Manitoulin Formation, Bruce Peninsula, Ontario

    Energy Technology Data Exchange (ETDEWEB)

    Anastas, A S; Coniglo, M [Waterloo Univ., ON (Canada)


    Several small, previously undescribed bioherms are present in the shallow shelf dolostones of the Manitoulin Formation at the Cabot Head and Wingfield Basin localities in the northernmost portion of the Bruce Peninsula region of southern Ontario. The bioherms, commonly associated with carbonate tempestites, range from 0.3 to 1.0 m in height and 0.9 to 2.5 m in width and are composed of bafflestones-floatstones and minor bindstones. The chief components of the bioherms are dolomitized lime mud and branching bryozoans. Bioherm building by bryozoans, although common in the ancient record, represents a great divergence from the mostly accessory frame encrusting role of bryozoans in modern environments. Minor skeletal components of the bioherms include echinoderms, rugose and tabulate corals and brachiopods. Laminar encrusting bryozoans exist in the top 10 cm of one of the bioherms. Some of the bioherms show evidence of water agitation that may be the result of current action induced by storm or tidal processes. The occurrence of the bioherms stretches the already known Llandoverian reef complex on Manitoulin Island further to the south. The reason why these bioherms did not reach sizes comparable to large Llandoverian or Wenlockian reefs and did not make the shift to coral-stromaporoid community is probably related to a complex interaction of factors such as community development, bathymetry, clasticity and salinity. 41 refs., 4 figs.

  8. Paleocommunity turnover in an Early Pliocene seamount from southeastern Spain (United States)

    García-Ramos, Diego Antonio; Zuschin, Martin


    Seamounts are topographic elevations under the sea, regardless of their size and relief. They support rich living communities and are important biodiversity hotspots, but many of the fundamental ecological processes that maintain seamount communities remain poorly understood. In contrast to snapshot observations conducted on extant seamounts, fossil examples may provide the opportunity to assess how temporal changes in physico-chemical parameters relate to paleocommunity turnovers in these particular biotopes. Here we deal with an Early Pliocene (Zanclean) small seamount in southeastern Spain. This classic locality is extremely rich in fossil macroinvertebrates and was subject to studies of some taxonomic groups in the late seventies. However, the detailed stratigraphy is herein outlined for the first time. The overall feature is a shallowing upward succession about 35 m thick which onlaps a Miocene volcanic ridge. The occurrence of the planktonic foraminifera Globorotalia margaritae and G. puncticulata allow attribution to the MPl3 biozone of the Mediterranean Pliocene. We measured two sections that can be divided in a lower interval of fine-grained bryozoan-rich deposits and a upper interval of biocalcarenite increasingly rich in rhodoliths upsection. The whole series is bioturbated, with Thalassinoides traces being more common upsection. Biofabrics comprise mostly densely-packed suites of disarticulated and fragmented shells of calcitic fauna (large oysters are often bioeroded by clionid sponges), suggesting relatively low sedimentation rates and reworking by storms (e.g., channelized shell-beds, tubular tempestites). The prevailing taxonomic groups are cheilostome bryozoans, oysters, brachiopods, pectinids, echinoderms, cirripedes and corals. The lower interval contains octocoral internodes (Isididae) (only recorded at the base of the section). Scleratinians like Balanophyllia? decrease in abundance upsection. Bryozoans are extremely abundant and diverse, with

  9. Biostratigraphy and biofacies of the Devonian deposits of the Kuh-e-Shorab section (southwest Damghan based on conodont fauna

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    Mehdi Hoveida


    Full Text Available The studied section located in the Djam-Semnan area 55 kms south west Damghan, at the vicinity of the Amrovan rail way station, in the northern flank of Shorab unassymetric anticline. The section is 440 meters thick, including limestone, dolomitic limestone and sandstone with rich fossils contents (e.g.: trilobite, coral, brachiopod, crinoid segments and vertebrate remains. The lower boundary is covered with the recent alluvial and the upper boundary disconformably overlain by the red marls and sandstones of the Permian Dorud formation. In order to determine biostratigraphy and bifacies of the deposits of Bahram formation in Shorab section, twenty five samples (4-6 kgs were collected and processed with conventional actic/formic acid. Nineteen samples were prolific and totally yielded 340 conodont elements. The extracted conodont elements mainly separated out of the carbonate units. Thirty-one species belong to four genera (Polygnathus, Icriodus, Pelecysgnathus and Ancyrodella were discriminated as: Icriodus excavatus, I. expansus, I. cedarensis, I. subterminus, I. iowaensis, I. alternatus, I. tafilaltensis, I. brevis, I. cf. expansus, I. sp., Polygnathus brevilaminus, Poly. angustidiscus, Poly. pollocki, Poly. cf. webbi, Poly. aspelundi, Poly. politus, Poly. alatus, Poly. webbi, Poly. cf. olgae, Poly. dubius, Poly. xylus, Poly. zinaidae, Poly. sp., Pelekeygnathus inclinathus, Pele. serradentatus, Pele. sp., Ancyrodella pristina, Acny. cf. pristina, Ancy. aff. binodosa, Ancy. sp. Based on the frequency and distribution of the collected elements Late Givetian to Early Famennian age were suggested to the studied interval. Conodonts biofacies (icriodid-polygnathid confirm the deposition of the Bahram formation of the Shorab section in the shallow carbonate condition.

  10. Climate change and the marine ecosystem of the western Antarctic Peninsula (United States)

    Clarke, Andrew; Murphy, Eugene J; Meredith, Michael P; King, John C; Peck, Lloyd S; Barnes, David K.A; Smith, Raymond C


    The Antarctic Peninsula is experiencing one of the fastest rates of regional climate change on Earth, resulting in the collapse of ice shelves, the retreat of glaciers and the exposure of new terrestrial habitat. In the nearby oceanic system, winter sea ice in the Bellingshausen and Amundsen seas has decreased in extent by 10% per decade, and shortened in seasonal duration. Surface waters have warmed by more than 1 K since the 1950s, and the Circumpolar Deep Water (CDW) of the Antarctic Circumpolar Current has also warmed. Of the changes observed in the marine ecosystem of the western Antarctic Peninsula (WAP) region to date, alterations in winter sea ice dynamics are the most likely to have had a direct impact on the marine fauna, principally through shifts in the extent and timing of habitat for ice-associated biota. Warming of seawater at depths below ca 100 m has yet to reach the levels that are biologically significant. Continued warming, or a change in the frequency of the flooding of CDW onto the WAP continental shelf may, however, induce sublethal effects that influence ecological interactions and hence food-web operation. The best evidence for recent changes in the ecosystem may come from organisms which record aspects of their population dynamics in their skeleton (such as molluscs or brachiopods) or where ecological interactions are preserved (such as in encrusting biota of hard substrata). In addition, a southwards shift of marine isotherms may induce a parallel migration of some taxa similar to that observed on land. The complexity of the Southern Ocean food web and the nonlinear nature of many interactions mean that predictions based on short-term studies of a small number of species are likely to be misleading. PMID:17405211

  11. A safer, urea-based in situ hybridization method improves detection of gene expression in diverse animal species. (United States)

    Sinigaglia, Chiara; Thiel, Daniel; Hejnol, Andreas; Houliston, Evelyn; Leclère, Lucas


    In situ hybridization is a widely employed technique allowing spatial visualization of gene expression in fixed specimens. It has greatly advanced our understanding of biological processes, including developmental regulation. In situ protocols are today routinely followed in numerous laboratories, and although details might change, they all include a hybridization step, where specific antisense RNA or DNA probes anneal to the target nucleic acid sequence. This step is generally carried out at high temperatures and in a denaturing solution, called hybridization buffer, commonly containing 50% (v/v) formamide - a hazardous chemical. When applied to the soft-bodied hydrozoan medusa Clytia hemisphaerica, we found that this traditional hybridization approach was not fully satisfactory, causing extensive deterioration of morphology and tissue texture which compromised our observation and interpretation of results. We thus tested alternative solutions for in situ detection of gene expression and, inspired by optimized protocols for Northern and Southern blot analysis, we substituted the 50% formamide with an equal volume of 8M urea solution in the hybridization buffer. Our new protocol not only yielded better morphologies and tissue consistency, but also notably improved the resolution of the signal, allowing more precise localization of gene expression and reducing aspecific staining associated with problematic areas. Given the improved results and reduced manipulation risks, we tested the urea protocol on other metazoans, two brachiopod species (Novocrania anomala and Terebratalia transversa) and the priapulid worm Priapulus caudatus, obtaining a similar reduction of aspecific probe binding. Overall, substitution of formamide by urea during in situ hybridization offers a safer alternative, potentially of widespread use in research, medical and teaching contexts. We encourage other workers to test this approach on their study organisms, and hope that they will also

  12. Stratigraphy and petroleum possibilities of lower Upper Devonian (Frasnian and lower Framennian) strata, Southwestern Utah (United States)

    Biller, Edward J.


    The lower Upper Devonian rocks in southwestern Utah--the Guilmette Formation and equivalents--represent a final regressive pulse of the major Late Devonian marine inundation of the Western Interior of the United States and record marine carbonate deposition on a wide continental shelf. They consist primarily of limestone, dolomite, and quartz arenite deposited in a shallow north-trending miogeosyncline, which constituted a single major basin of accumulation on this shelf. The Guilmette Formation and equivalents were deposited in shallow normal to hypersaline marine waters. The environments of deposition include: a moderate- to high-energy intertidal environment, a moderate-energy subtidal environment, a lower energy, deeper subtidal environment below effective wave base, and a high-energy environment in local shallow areas of mud mounds and bioherms. The carbonate deposition of the Guilmette Formation and equivalents was interrupted periodically by the deposition of quartz arenites. These may represent the breaking up of the miogeosynclinal-cratonic pattern of deposition. In most areas, the Guilmette and equivalents are overlain by a thin transgressive marine quartz arenite deposit--the Cove Fort Quartzite and basal Leatham equivalent. Previous paleontologic evidence indicated a general Middle to Late Devonian age for the Guilmette Formation. The present study narrows this range and suggests that the age of the Guilmette Formation and its equivalents is late Middle Devonian (Stringocephalus brachiopod zone) to early Late Devonian (Uppermost Palmatolepis gigas conodont zone). Available subsurface data suggest that the petroleum possibilities of the Guilmette Formation and equivalents in southwestern Utah are poor. Several tests have penetrated .the interval with only minor shows of oil in rocks with low porosity and permeability. Nevertheless, many outcrop samples of the same interval, appear to have excellent porosity and permeability and a strongly fetid odor,

  13. The importance of Hindeodus parvus (Conodonta for the definition of the Permian-Triassic boundary and evaluation of the proposed sections for a global stratotype section and point (GSSP for the base of the Triassic

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    Yurij D. Zakharov


    Full Text Available The biostratigraphic Permian/Triassic (P/T boundary is defined by the first appearance of H. parvus. The first appearance of H. parvus within the dine H. latidentatus-H. parvus is a globally recognizable event in the conodont evolution.The first appearance of H. parvus is not facies related and can be observed both in ammonoid-free shallow-water deposits and in ammonoid-bearing pelagic deposits. H. parvus is a common, easily determinable species known so far from the entire Tethys, Japan, western North America, Boreal realm (Greenlandand the Tethyan margin of Gondwana. H. parvus is the first species withworld-wide distribution to appear after the absolute minimum in the faunal diversity indicated by the minimum in The Meishan section (South China contains a continuous, pelagic sedimentaryrecord across the P/T boundary without stratigraphie gaps. It is nearly unaltered thermally (CAI = 1-1.5. Its fossil content (ammonoids, conodonts, foraminifers,bivalves, brachiopods, sporomorphs etc. and event succession have been thoroughlystudied. Absolute age and magnetostratigraphy have also been subjected to intensive studies. The section is readily accessible and under protection of the government. This section is best suitable as a global stratotype section and point (GSSP for the base of the Triassic. No other section in the world is known tobe qualified for defining the P/T boundary in a GSSP. H. parvus made its earliest appearence in the middle part of Boundary Bed 2 (Bed 27 at Meishan. It evolved within Bed 27 from H. latidentatus within a phylomorphogenetic continuum in a continuous and monofacial stratum. The biostratigraphic P/T boundary lies very close to the event boundary (15 cm above the event boundary at the baseof Boundary Bed 1 = Bed 25, and a few centimetres above the minimum in Ô'13C in the lower Boundary Bed 2.

  14. Phanerozoic trends in shell accumulations: A comparison of Ordovician-Silurian and Tertiary records

    Energy Technology Data Exchange (ETDEWEB)

    Kidwell, S.M. (Univ. of Chicago, IL (United States). Dept. of Geophysical Sciences); Brenchley, P.J. (Univ. of Liverpool (United Kingdom). Dept. of Earth Sciences)


    Evolutionary changes in the diversity and bathymetric deployment of major benthic invertebrate groups, both those that produce and those that modify bioclasts, suggest that the nature of shell accumulations is likely to have also changed through the Phanerozoic. Specifically, it has been suggested that, excepting crinoidal deposits, the Paleozoic-Triassic record is dominated by thin shell concentrations (< few 10s cm) whereas the Cretaceous-Cenozoic record includes much thicker accumulations (> 1 m). To test this, the authors focused on siliciclastic records in two end-point periods. Using the literature on North America and Europe combined with extensive field-checking, they found that brachiopod-, trilobite-, and bryozoan-dominated accumulations in the Ord-Sil are typically thin but that there are important exceptions. In the Tertiary, thick concentrations are composed of a much larger variety of groups, occur in a wider range of environments, and exhibit a greater diversity of biostratinomic features and inferred accumulation histories. Information on the paleobathymetry, paleolatitude and tectonic setting was evaluated for all data points, and so the contrasting patterns of shell accumulation can clearly be related to differences in geologic age rather than to environmental biases in data collection. Even the analysis of only a few shell-bed features, therefore, provides strong evidence for long-term changes in the nature of the bioclastic fossil record. Work still in progress suggests that there are also long-term changes in the distribution and nature of bioclastic deposits through stratigraphic (3rd-order) sequences, particularly in the character of skeletal material associated with key surfaces.

  15. Missourian (early Late Pennsylvanian) climate in Midcontinent North America

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    Schutter, S.R.; Heckel, P.H.


    The abrupt decrease in mineable coals from Desmoinesian to Missourian rocks in Midcontinent North America has been related by several lines of evidence to the probability that Missourian climate became at least seasonally drier than Desmoinesian climate. This represents a transition from the equatorial Desmoinesian rainforest climate to the arid Permian climate, as North America moved northward from the equator. This change is reflected in the progression of evaporites from western Colorado in the Desmoinesian to Kansas in the Permian. Direct climatic evidence from soils in two Missourian shales deposited at low stands of sea-level includes caliche horizons, incompletely leached mixed-layer clays. Less direct climatic evidence includes the greater proportion of marine limestone deposited at intermediate sea-level stands in the Missourian than in the Desmoinesian part of the sequence, in conjunction with the greater abundance and thickness of oolite and shore-line facies in Missourian limestones than in Desmoinesian counterparts. This probably reflects increasing dryness of the climate, which would have led to decreased detrital influx and increased salinity as rainfall and runoff diminished. Indirect climatic evidence in offshore black phosphatic shales deposited at highest sea-level stands involves possible seasonality in organic and phosphatic laminations, related to periodicity of upwelling in the tropical trade-wide belt under the strong monsoonal influence of Pangaea. Mid-continent Missourian climates probably ranged from tropical monsoon to tropical savanna or hot steppe, all having wet-dry seasonality. The coincident extinctions of conodont, brachiopod, and other marine genera as well as the loss of swamp lycopods at the end of the Desmoinesian suggest the possibility that a greater than usual drop in sea level affected both realms at this time, while the climate was becoming drier. 91 references.


    International Nuclear Information System (INIS)

    Deines, Steven D.; Williams, Carol A.


    This paper determines Earth's rotational deceleration without relying on atomic or ephemeris timescales. Earth's rotation defines the civil time standard called Universal Time (UT). Our previous paper did not examine tidal friction in depth when analyzing the timescale divergence between UT and International Atomic Time (TAI). We examine all available paleontological fossils and deposits for the direct measurements of Earth's past rotation rates, because that record includes all contributing effects. We examine paleontological reports that date Earth's rotation rate using corals, bivalves, brachiopods, rhythmites, and stromatolites. Contributions that vary Earth's moment of inertia, such as continental plate drifts, coastline changes, ice age formations, and viscous glacial rebounds, are superimposed with the secular deceleration. The average deceleration of Earth's rotation rate from all available fossil data is found to be (5.969 ± 1.762) × 10 −7 rad yr −2 . Our value is 99.8% of the total rotational deceleration determined by Christodoulidis et al., who used artificial satellite data, and our value is 96.6% of the expected tidal friction value obtained by Stephenson and Morrison. Taking the derivative of conserved angular momentum, the predicted lunar orbital deceleration caused by the average rotational deceleration corresponds closely to lunar models. When evaluating the significant time gaps between UT and TAI, Earth's rotational deceleration is a minor contributing factor. Also, the secular deceleration rate is necessary to correctly date ancient astronomical events. We strongly encourage that more ocean paleontological evidence be found to supplement the record to separate the many periodic variations embedded in these data

  17. An evaluation of intertidal feeding habitats from a shorebird perspective: Towards relevant comparisons between temperate and tropical mudflats (United States)

    Piersma, Theunis; de Goeij, Petra; Tulp, Ingrid

    Macrozoobenthic communities of intertidal soft sediments are reviewed worldwide from the perspective of a mollusc-eating shorebird species. Based on 19 sites, total biomass figures varied between 5 and 80 g AFDM per m 2 (average 24 g AFDM per m 2); no latitudinal trends are apparent. The contribution made by bivalves and gastropods varies between 1% and 99%, north-temperate intertidal flats having relatively more molluscs than tropical flats. Intertidal flats in the tropics contain a greater variety of taxa, with brachiopods in Indonesia and echinoderms in northwest Australia contributing significantly to biomass only there. Limits to the occurrence of avian predators of intertidal benthos are set by the harvestable fraction of the biomass on offer and the costs of living at a particular site. No systematic differences in the harvestable fraction of the total mollusc-biomass for a worldwide occurring shorebird species specializing on molluscs (knots Calidris canutus) were apparent between temperate and tropical intertidal areas, in spite of large differences in maintenance metabolism incurred by these birds. The harvestable fractions of bivalves in the two West African areas (Banc d'Arguin, Mauritania and Guinea-Bissau) tended to be high (23-84% of total biomass in six species), they were relatively low (2-52% in five species) in the temperate Wadden Sea and the tropical northwest Australian site. Harvestable biomass determines the intake rate of shorebirds, as illustrated by functional-response curves of knots feeding on two bivalves species. We argue that the collection of information on size-depth relationships along with faunal and biomass surveys at a range of sites is bound to greatly increase our understanding of both the biology of tidal-flat invertebrates and the resource base underpinning the spectacular seasonal migrations of shorebirds.

  18. Early Silurian (Llandoverian) Leask Point and Charlton Bay bioherms, Manitoulin Island, Ontario, Canada

    Energy Technology Data Exchange (ETDEWEB)

    Mielczarek, W.; Copper, P.


    About 300 bioherms are known in the Llandoverian Manitoulin Formation of eastern Manitoulin Island. In the South Bay area, the large Leask Piont bioherm and Charlton Bay patch-reef complex lack a distinct skeletal growth framework. Bioherms consist of mudstone and wackestone, with isolated lenses of bafflestone, boundstone, floatstone. Fossils are scarce, but crinozoans and bryozoans comprise about 90% of the bioclasts. Other fauna include stromatoporoids, corals, brachiopods, gastropods, trilobites, and probable algae (algae are difficult to identify and may have played a significant role). Faunal ratios remained relatively constant during mound growth. Soft substrates with sedimentation rates of a few millimeters per year are suggested by bedding type and morphologic dominance of lamellar and tabular corals and stromatoporoids. An increased sedimentation rate, resulting from shoaling, is indicated by more overturned, broadly conical corals in the upper parts of the mounds. Shoaling may be responsible for cessation of mound growth. Lithoclasts are more common in the upper parts of the mounds. They formed when semiconsolidated muds were disturbed and redeposited during storms. Megarippled interreef surface areas, largely devoid of coral growth, indicate mud instability at Charlton Bay. Lack of suitable stable substrates may have hampered coral development. Dolomitization was postdepositional. The diagenetic sequence occurred in three stages: 1)selective pyritization and silicification, formation of an early muddy dolomite replacing the mud fraction of the dolostone, lithification and formation of rare calcite cement and neomorphic syntaxial rims; 2)clear, coarse dolomite replacing pore-filling calcite cement, syntaxial rims, and unaltered macrofossils, stylolitization, grain-to-grain dissolution; and 3)a late dolomite found mainly as fine rhombs in stylolites, solution seams, and intraskeletal pore space.

  19. El Paso Formation - a Lower Ordovician platform carbonate deposit

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    Clemons, R.E.


    The eastward-transgressive Lower Ordovician El Paso Formation conformably overlies Bliss Sandstone in southern New Mexico. Locally, lower El Paso was deposited on low hills of plutonic and volcanic rocks. The region subsided gradually throughout Canadian time, receiving the El Paso carbonate rock blanket up to 460 m thick. Lithologic and chronologic correlative rocks were deposited over most of the southwestern US as the first Paleozoic carbonate platform sequence. The El Paso Formation contains four members, listed here in ascending order: Hitt Canyon, Jose, McKelligon, and Padre. Gradually decreasing sand content upward through the Hitt Canyon indicates deepening water and/or greater distance to shore. Girvanella(.) oncolites are locally abundant. Stromatolite mounds near the top of the Hitt Canyon, combined with an influx of sand, ooids, and rounded bioclasts in the Jose Member, recorded a shoaling phase. The overlying McKelligon Member contains little or no sand, and sponge-Calathium mounds are prominent at some locales. Stromatolite mounds are interbedded with sponge-Calathium mounds in a few sections. Lower Padre Member beds are typically silty to sandy and locally contain thinly-laminated zones. The Padre contains more restricted fauna that includes traces of ostracods. Pervasive bioturbation of El Paso beds and fauna consisting of echinoderms, sponges, gastropods, trilobites, Nuia, Calathium, cephalopods, and algae plus minor brachiopods and Pulchrilamina indicate predominating shallow-subtidal environments. Low-energy platform environments, in which a large volume of micritic muds accumulated, were disturbed thousands of times by storms producing abundant thin, poorly washed biosparite, intrasparite, and intrasparrudite lenses.

  20. Simpson-Arbuckle contact revisited in Northwest Oklahoma County, Oklahoma

    Energy Technology Data Exchange (ETDEWEB)

    Allison, M.D.; Allen, R.W. [Kabodi Inc., Ardmore, OK (United States)


    The Joins Formation, the lowermost formation of the Simpson Group, is traditionally the least studied or understood of the Simpson formations. The Joins, not known to produce hydrocarbons in central Oklahoma, is frequently overlooked by those more interested in the productive Simpson formations above and the Arbuckle carbonates below. In a study of the lower Simpson to upper Arbuckle interval in northwestern Oklahoma County, Oklahoma, the Joins Formation was found to be present. The central Oklahoma section consists of interbedded gray, olive gray and green splintery moderately waxy shale, cream to light gray homogeneous microcrystallin dolomite, and microcrystalline to fine crystalline fossiliferous slightly glauconitic well cemented sandstones are also noted. The entire Joins Formation is moderately to very fossiliferous; primarily consisting of crinoids, ostracods, brachiopods, and trilobites. The ostracod fauna closely resembles and correlates with the Arbuckle Mountain section, which has been extensively studied over the years by such authors as Taff, Ulrich and Harris. Beneath the Joins in this area is a normal section of Arbuckle dolomites. Due to the absence of a basal sand in the Joins the separation of the Joins and Arbuckle, utilizing electric logs only, is frequently tenuous. In comparison with the Arbuckle, the Joins tends to have higher gamma ray and S.P. values. Other tools, such as resistivity, bulk density and photoelectric (PE), are frequently inconclusive. For geologists studying the Simpson-Arbuckle contact in central Oklahoma, the presence or absence of the Joins Formation is best determined through conventional lithologic and palenontologic sample identification techniques. Once this has been done, correlation of electric logs with this type log is possible for the local area.

  1. Lower Paleozoic carbonate rocks of Baird Mountains Quadrangle, Alaska

    Energy Technology Data Exchange (ETDEWEB)

    Dumoulin, J.A.; Harris, A.G.


    Lower Paleozoic carbonate rocks in the Baird Mountains quadrangle form a relatively thin (about 550 m), chiefly shallow-water succession that has been imbricately thrust and metamorphosed to lower greenschist facies. Middle and Upper Cambrian rocks - the first reported from the western Brooks Range - occur in the northeastern quarter of the quadrangle, south of Angayukaqsraq (formerly Hub) Mountain. They consist of marble grading upward into thin-bedded marble/dolostone couplets and contain pelagiellid mollusks, acetretid brachiopods, and agnostid trilobites. Sedimentologic features and the Pelagiellas indicate a shallow-water depositional environment. Overlying these rocks are Lower and Middle Ordovician marble and phyllite containing graptolites and conodonts of midshelf to basinal aspect. Upper Ordovician rocks in this area are bioturbated to laminated dolostone containing warm, shallow-water conodonts. In the Omar and Squirrel Rivers areas to the west, the Lower Ordovician carbonate rocks show striking differences in lithofacies, biofacies, and thickness. Here they are mainly dolostone with locally well-developed fenestral fabric and evaporite molds, and bioturbated to laminated orange- and gray-weathering dolomitic marble. Upper Silurian dolostone, found near Angayukaqsraq Mountain and on the central Squirrel River, contains locally abundant corals and stronmatoporoids. Devonian carbonate rocks are widely distributed in the Baird Mountains quadrangle; at least two distinct sequences have been identified. In the Omar area, Lower and Middle Devonian dolostone and marble are locally cherty and rich in megafossils. In the north-central (Nakolik River) area, Middle and Upper Devonian marble is interlayered with planar to cross-laminated quartz-carbonate metasandstone and phyllite.

  2. Geologic drivers of late ordovician faunal change in laurentia: investigating links between tectonics, speciation, and biotic invasions.

    Directory of Open Access Journals (Sweden)

    David F Wright

    Full Text Available Geologic process, including tectonics and global climate change, profoundly impact the evolution of life because they have the propensity to facilitate episodes of biogeographic differentiation and influence patterns of speciation. We investigate causal links between a dramatic faunal turnover and two dominant geologic processes operating within Laurentia during the Late Ordovician: the Taconian Orogeny and GICE related global cooling. We utilize a novel approach for elucidating the relationship between biotic and geologic changes using a time-stratigraphic, species-level evolutionary framework for articulated brachiopods from North America. Phylogenetic biogeographic analyses indicate a fundamental shift in speciation mode-from a vicariance to dispersal dominated macroevolutionary regime-across the boundary between the Sandbian to Katian Stages. This boundary also corresponds to the onset of renewed intensification of tectonic activity and mountain building, the development of an upwelling zone that introduced cool, nutrient-rich waters into the epieric seas of eastern Laurentia, and the GICE isotopic excursion. The synchronicity of these dramatic geologic, oceanographic, and macroevolutionary changes supports the influence of geologic events on biological evolution. Together, the renewed tectonic activity and oceanographic changes facilitated fundamental changes in habitat structure in eastern North America that reduced opportunities for isolation and vicariance. They also facilitated regional biotic dispersal of taxa that led to the subsequent establishment of extrabasinal (=invasive species and may have led to a suppression of speciation within Laurentian faunas. Phylogenetic biogeographic analysis further indicates that the Richmondian Invasion was a multidirectional regional invasion event that involved taxa immigrating into the Cincinnati region from basins located near the continental margins and within the continental interior.

  3. The Jurassic of Denmark and Greenland: The Upper Jurassic of Europe: its subdivision and correlation

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    Zeiss, Arnold


    Full Text Available In the last 40 years, the stratigraphy of the Upper Jurassic of Europe has received much attention and considerable revision; much of the impetus behind this endeavour has stemmed from the work of the International Subcommission on Jurassic Stratigraphy. The Upper Jurassic Series consists of three stages, the Oxfordian, Kimmeridgian and Tithonian which are further subdivided into substages, zones and subzones, primarily on the basis of ammonites. Regional variations between the Mediterranean, Submediterranean and Subboreal provinces are discussed and correlation possibilities indicated. The durations of the Oxfordian, Kimmeridgian and Tithonian Stages are reported to have been 5.3, 3.4 and 6.5 Ma, respectively. This review of the present status of Upper Jurassic stratigraphy aids identification of a number of problems of subdivision and definition of Upper Jurassic stages; in particular these include correlation of the base of the Kimmeridgian and the top of the Tithonian between Submediterranean and Subboreal Europe. Although still primarily based on ammonite stratigraphy, subdivision of the Upper Jurassic is increasingly being refined by the incorporation of other fossil groups; these include both megafossils, such as aptychi, belemnites, bivalves, gastropods, brachiopods, echinoderms, corals, sponges and vertebrates, and microfossils such as foraminifera, radiolaria, ciliata, ostracodes, dinoflagellates, calcareous nannofossils, charophyaceae, dasycladaceae, spores and pollen. Important future developments will depend on the detailed integration of these disparate biostratigraphic data and their precise combination with the abundant new data from sequence stratigraphy, utilising the high degree of stratigraphic resolution offered by certain groups of fossils. This article also contains some notes on the recent results of magnetostratigraphy and sequence chronostratigraphy.

  4. Preliminary report on the Oldenburg “butter shale” in the Upper Ordovician (Katian; Richmondian Waynesville Formation, USA

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    Christopher D. Aucoin


    Full Text Available The Cincinnatian Series (Upper Ordovician; upper Katian of the Cincinnati Arch region, Ohio, Indiana and Kentucky contains several bed packages informally referred to as “butter shales” or “trilobite shales”. These packages are typically 1–2 m of relatively pure, homogeneous claystone with isolated, lenticular limestone beds. These claystones are most widely known for their excellent preservation of abundant trilobites, especially Isotelus and Flexicalymene, as well as diverse and commonly articulated bivalves, and nautiloids. A newly recognized butter shale interval in the Clarksville Member of the Waynesville Formation contains a typical butter-shale fossil assemblage, dominated by bivalves, orthoconic cephalopods and trilobites. To better study the fabric of this claystone, a large, epoxy-coated block of the claystone was dry-cut. Polished surfaces show a variety of otherwise cryptic features, including pervasive bioturbation and the presence of probable lingulid escape burrows (Lingulichnus, as well as abundant fodinichnia (Chondrites, Planolites, Teichichnus. Preservation of articulated trilobites and closed bivalves in approximate living position, as well as escape burrows, indicates deposition as a series of mud burial events or obrution deposits. We suggest that the butter shales resulted from net accumulation of multiple episodes of re-suspended mud deposition, which rapidly smothered organisms and resulted in exceptional preservation. Between events the seafloor was colonized by abundant deposit-feeding infaunal organisms, which destabilized the substrate and generated turbidity near the sediment–water interface, thus inhibiting sessile suspension feeders. Rapid net deposition was also interrupted by more prolonged periods (tens to hundreds of years of low sedimentation that permitted colonization by epifaunal brachiopod-dominated communities. While most butter shale units are regionally extensive, the Oldenburg is

  5. Calibrating water depths of Ordovician communities: lithological and ecological controls on depositional gradients in Upper Ordovician strata of southern Ohio and north-central Kentucky, USA

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    Carlton E. Brett


    Full Text Available Limestone and shale facies of the Upper Ordovician Grant Lake Formation (Katian: Cincinnatian, Maysvillian are well exposed in the Cincinnati Arch region of southern Ohio and north-central Kentucky, USA. These rocks record a gradual change in lithofacies and biofacies along a gently northward-sloping ramp. This gradient spans very shallow, olive-gray, platy, laminated dolostones with sparse ostracodes in the south to offshore, nodular, phosphatic, brachiopod-rich limestones and marls in the north. This study uses facies analysis in outcrop to determine paleoenvironmental parameters, particularly those related to water depth (e.g., position of the photic zone and shoreline, relative degree of environmental energy. Within a tightly correlated stratigraphic interval (the Mount Auburn and Straight Creek members of the Grant Lake Formation and the Terrill Member of the Ashlock Formation, we document the occurrence of paleoenvironmental indicators, including desiccation cracks and light-depth indicators, such as red and green algal fossils and oncolites. This permitted recognition of a ramp with an average gradient of 10–20 cm water depth per horizontal kilometer. Thus, shallow subtidal (“lagoonal” deposits in the upramp portion fall within the 1.5–6 m depth range, cross-bedded grainstones representing shoal-type environments fall within the 6–18 m depth range and subtidal, shell-rich deposits in the downramp portion fall within the 20–30 m depth range. These estimates match interpretations of depth independently derived from faunal and sedimentologic evidence that previously suggested a gentle ramp gradient and contribute to ongoing and future high-resolution paleontologic and stratigraphic studies of the Cincinnati Arch region.


    Energy Technology Data Exchange (ETDEWEB)

    Deines, Steven D. [Donatech Corporation, Fairfield, IA 52556 (United States); Williams, Carol A., E-mail:, E-mail: [Department of Mathematics and Statistics (Prof. emeritus), University of South Florida, 4202 E. Fowler Ave., Tampa, FL 33620 (United States)


    This paper determines Earth's rotational deceleration without relying on atomic or ephemeris timescales. Earth's rotation defines the civil time standard called Universal Time (UT). Our previous paper did not examine tidal friction in depth when analyzing the timescale divergence between UT and International Atomic Time (TAI). We examine all available paleontological fossils and deposits for the direct measurements of Earth's past rotation rates, because that record includes all contributing effects. We examine paleontological reports that date Earth's rotation rate using corals, bivalves, brachiopods, rhythmites, and stromatolites. Contributions that vary Earth's moment of inertia, such as continental plate drifts, coastline changes, ice age formations, and viscous glacial rebounds, are superimposed with the secular deceleration. The average deceleration of Earth's rotation rate from all available fossil data is found to be (5.969 ± 1.762) × 10{sup −7} rad yr{sup −2}. Our value is 99.8% of the total rotational deceleration determined by Christodoulidis et al., who used artificial satellite data, and our value is 96.6% of the expected tidal friction value obtained by Stephenson and Morrison. Taking the derivative of conserved angular momentum, the predicted lunar orbital deceleration caused by the average rotational deceleration corresponds closely to lunar models. When evaluating the significant time gaps between UT and TAI, Earth's rotational deceleration is a minor contributing factor. Also, the secular deceleration rate is necessary to correctly date ancient astronomical events. We strongly encourage that more ocean paleontological evidence be found to supplement the record to separate the many periodic variations embedded in these data.

  7. Trilobite-based biostratigraphy (arthropoda-trilobita) and related faunas of the Cambrian from Sonora, Mexico (United States)

    Cuen-Romero, F. J.; Valdez-Holguín, J. E.; Buitrón-Sánchez, B. E.; Monreal, R.; Enríquez-Ocaña, L. F.; Aguirre-Hinojosa, E.; Ochoa-Granillo, J. A.; Palafox-Reyes, J. J.


    A biostratigraphic analysis based on trilobites of the main Cambrian outcrops from Sonora, Mexico is performed. The data are based on a combination of field work and published sources, including four previously studied locations from northern and eastern Sonora (Caborca, Cananea, Mazatán, and Arivechi) as well as a new location in the central part of the state of Sonora (San José de Gracia). Chronostratigraphic positions are assigned to the Cambrian outcrops according to Peng et al., 2012 and Cohen et al., 2017. In the Caborca area, the Puerto Blanco, Proveedora, Buelna, Cerro Prieto, Arrojos and El Tren formations comprise a wide range of biozones, which starts from the Fritzaspis Zone until the Glossopleura walcotti Zone (Begadean-Lincolnian Series, global Stage 3-Stage 5, Series 2-Series 3). The Bolsa Quartzite and the Abrigo Limestone exposed in Cananea area are assigned to the Cedaria/Cedarina dakotaensis Zone until the Crepicephalus Zone (Lincolnian Series-Marjuman Stage, global Series 3-Guzhangian). In the San José de Gracia area, The Proveedora, Buelna, Cerro Prieto and El Gavilán formations range from the ?Bristolia mohavensis or ?Bristolia insolens zones until the upper part of Mexicella mexicana Zone, Albertella highlandensis Subzone (Series 2-Series 3, Stage 4-Stage 5). In the Arivechi area, the La Sata, El Mogallón, La Huerta and the Milpillas formations range from Poliella denticulata Zone to the Elvinia Zone (Lincolnian-Millardan, Delamaran-Steptoean, global Series 3-Furongian, Stage 5-Paibian). Paleozoic marine fauna distribution in northwest Mexico and the southwest United States of America, suggest that during this time an extensive faunal province existed, containing a great variety of marine invertebrates with notorious intraspecific affinity. The biotic association includes poriferans, archaeocyathids, brachiopods, mollusks, arthropods and echinoderms as predominant elements.

  8. Recovery vs. Restructuring: Establishing Ecologic Patterns in Early and Middle Triassic Paleocommunities (Invited) (United States)

    Fraiser, M.; Dineen, A.; Sheehan, P.


    Published data has been interpreted as indicating that marine ecological devastation following the end-Permian mass extinction was protracted and may have lasted 5 million years into the Middle Triassic (Anisian). However, a review of previous literature shows that understanding of biotic recovery is typically based on only a few components of the ecosystem, such as on taxonomic diversity, a single genus/phylum, or facies. Typically, paleocommunities are considered fully recovered when dominance and diversity are regained and normal ecosystem functioning has resumed. However, in addition to the biodiversity crash at the end of the Permian, taxonomic and ecologic structure also changed,with the extinction marking the faunal shift from brachiopod-rich Paleozoic Evolutionary Fauna (EF) to the mollusc-rich Modern EF. This suggests that the extreme reorganizational nature of the Triassic does not adhere to the standard definition of recovery, which is a return to previous conditions. Thus, we propose the term 'restructuring' to describe this interval, as Early and Middle Triassic communities might not exhibit the typical characteristics of a 'normal' Permian one. To more fully characterize Triassic ecologic restructuring, paleoecologists should take into account functional diversity and redundancy. We quantified functional richness and regularity in four different paleocommunities from classic Permian and Triassic sections. Functional richness was low in paleocommunities after the end-Permian mass extinction, but increased to high levels by the Middle Triassic. In contrast, functional regularity was low in the Middle Permian, but high in all the Triassic paleocommunities. The change from low to high functional regularity/redundancy at the P/T boundary may be a factor of the highly stressful Triassic environmental conditions (i.e. anoxia, hypercapnia), as high regularity in a community can boost survival in harsh environments. Parameters such as these will more

  9. Temperature evolution and the oxygen isotope composition of Phanerozoic oceans from carbonate clumped isotope thermometry (United States)

    Henkes, Gregory A.; Passey, Benjamin H.; Grossman, Ethan L.; Shenton, Brock J.; Yancey, Thomas E.; Pérez-Huerta, Alberto


    Surface temperature is among the most important parameters describing planetary climate and habitability, and yet there remains considerable debate about the temperature evolution of the Earth's oceans during the Phanerozoic Eon (541 million years ago to present), the time during which complex metazoan life radiated on Earth. Here we critically assess the emerging record of Phanerozoic ocean temperatures based on carbonate clumped isotope thermometry of fossil brachiopod and mollusk shells, and we present new data that fill important gaps in the Late Paleozoic record. We evaluate and reject the hypothesis that solid-state reordering of 13C-18O bonds has destroyed the primary clumped isotope temperature signal of most fossils during sedimentary burial at elevated temperatures. The resulting Phanerozoic record, which shows a general coupling between tropical seawater temperatures and atmospheric carbon dioxide (CO2) levels since the Paleozoic, indicates that tropical temperatures during the icehouse climate of the Carboniferous period were broadly similar to present (∼25-30 °C), and suggests that benthic metazoans were able to thrive at temperatures of 35-40 °C during intervals of the early and possibly the latest Paleozoic when CO2 levels were likely 5-10× higher than present-day values. Equally important, there is no resolvable trend in seawater oxygen isotope ratios (δ18 O) over the past ∼500 million years, indicating that the average temperature of oxygen exchange between seawater and the oceanic crust has been high (∼270 °C) since at least the early Paleozoic, which points to mid-ocean ridges as the dominant locus of water-rock interaction over the past half-billion years.

  10. Microconchids from microbialite ecosystem immediately after end-Permian mass extinction: ecologic selectivity and implications for microbialite ecosystem structure (United States)

    Yang, H.; Chen, Z.; Wang, Y. B.; Ou, W.; Liao, W.; Mei, X.


    The Permian-Triassic (P-Tr) carbonate successions are often characterized by the presence of microbialite buildups worldwide. The widespread microbialites are believed as indication of microbial proliferation immediately after the P-Tr mass extinction. The death of animals representing the primary consumer trophic structure of marine ecosystem in the P-Tr crisis allows the bloom of microbes as an important primary producer in marine trophic food web structure. Thus, the PTB microbialite builders have been regarded as disaster taxa of the P-Tr ecologic crisis. Microbialite ecosystems were suitable for most organisms to inhabit. However, increasing evidence show that microbialite dwellers are also considerably abundant and diverse, including mainly foraminifers Earlandia sp. and Rectocornuspira sp., lingulid brachiopods, ostrocods, gastropods, and microconchids. In particular, ostracods are extremely abundant in this special ecosystem. Microconchid-like calcareous tubes are also considerably abundant. Here, we have sampled systematically a PTB microbialite deposit from the Dajiang section, southern Guizhou Province, southwest China and have extracted abundant isolated specimens of calcareous worm tubes. Quantitative analysis enables to investigate stratigraphic and facies preferences of microconchids in the PTB microbialites. Our preliminary result indicates that three microconchid species Microconchus sp., Helicoconchus elongates and Microconchus aberrans inhabited in microbialite ecosystem. Most microconchilds occurred in the upper part of the microbialite buildup and the grainstone-packstone microfacies. Very few microconchilds were found in the rocks bearing well-developed microbialite structures. Their stratigraphic and environmental preferences indicate proliferation of those metazoan organisms is coupled with ebb of the microbialite development. They also proliferated in some local niches in which microbial activities were not very active even if those

  11. Climate change and the marine ecosystem of the western Antarctic Peninsula. (United States)

    Clarke, Andrew; Murphy, Eugene J; Meredith, Michael P; King, John C; Peck, Lloyd S; Barnes, David K A; Smith, Raymond C


    The Antarctic Peninsula is experiencing one of the fastest rates of regional climate change on Earth, resulting in the collapse of ice shelves, the retreat of glaciers and the exposure of new terrestrial habitat. In the nearby oceanic system, winter sea ice in the Bellingshausen and Amundsen seas has decreased in extent by 10% per decade, and shortened in seasonal duration. Surface waters have warmed by more than 1 K since the 1950s, and the Circumpolar Deep Water (CDW) of the Antarctic Circumpolar Current has also warmed. Of the changes observed in the marine ecosystem of the western Antarctic Peninsula (WAP) region to date, alterations in winter sea ice dynamics are the most likely to have had a direct impact on the marine fauna, principally through shifts in the extent and timing of habitat for ice-associated biota. Warming of seawater at depths below ca 100 m has yet to reach the levels that are biologically significant. Continued warming, or a change in the frequency of the flooding of CDW onto the WAP continental shelf may, however, induce sublethal effects that influence ecological interactions and hence food-web operation. The best evidence for recent changes in the ecosystem may come from organisms which record aspects of their population dynamics in their skeleton (such as molluscs or brachiopods) or where ecological interactions are preserved (such as in encrusting biota of hard substrata). In addition, a southwards shift of marine isotherms may induce a parallel migration of some taxa similar to that observed on land. The complexity of the Southern Ocean food web and the nonlinear nature of many interactions mean that predictions based on short-term studies of a small number of species are likely to be misleading.

  12. Reconstructing the diet of a 505-million-year-old arthropod: Sidneyia inexpectans from the Burgess Shale fauna. (United States)

    Zacaï, Axelle; Vannier, Jean; Lerosey-Aubril, Rudy


    The feeding ecology of the 505-million-year-old arthropod Sidneyia inexpectans from the middle Cambrian (Series 3, Stage 5) Burgess Shale fauna (British Columbia, Canada) is revealed by three lines of evidence: the structure of its digestive system, the fossilized contents of its gut and the functional anatomy of its appendages. The digestive tract of Sidneyia is straight, tubular and relatively narrow in the trunk region. It is enlarged into a pear-shaped area in the cephalic region and stretches notably to form a large pocket in the abdomen. The mouth is ventral, posteriorly directed and leads to the midgut via a short tubular structure interpreted as the oesophagus. Anteriorly, three pairs of glands with internal, branching tubular structures open into the digestive tract. These glands have equivalents in various Cambrian arthropod taxa (e.g. naraoiids) and modern arthropods. Their primary function was most likely to digest and assimilate food. The abdominal pocket of Sidneyia concentrates undigested skeletal elements and various residues. It is interpreted here as the functional analogue of the stercoral pocket of some extant terrestrial arachnids (e.g. Araneae, Solifugae), whose primary function is to store food residuals and excretory material until defecation. Analysis of the gut contents indicates that Sidneyia fed largely on small ptychopariid trilobites, brachiopods, possibly agnostids, worms and other undetermined animals. Sidneyia was primarily a durophagous carnivore with predatory and/or scavenging habits, feeding on small invertebrates that lived at the water-sediment interface. There is no evidence for selective feeding. Its food items (e.g. living prey or dead material) were grasped and manipulated ventrally by its anterior appendages, then macerated into ingestible fragments and conveyed to the mouth via the converging action of strong molar-like gnathobases. Digestion probably took place within the anterior midgut via enzymes secreted in the

  13. Complete mitochondrial genome of Bugula neritina (Bryozoa, Gymnolaemata, Cheilostomata: phylogenetic position of Bryozoa and phylogeny of lophophorates within the Lophotrochozoa

    Directory of Open Access Journals (Sweden)

    Jang Kuem


    Full Text Available Abstract Background The phylogenetic position of Bryozoa is one of the most controversial issues in metazoan phylogeny. In an attempt to address this issue, the first bryozoan mitochondrial genome from Flustrellidra hispida (Gymnolaemata, Ctenostomata was recently sequenced and characterized. Unfortunately, it has extensive gene translocation and extremely reduced size. In addition, the phylogenies obtained from the result were conflicting, so they failed to assign a reliable phylogenetic position to Bryozoa or to clarify lophophorate phylogeny. Thus, it is necessary to characterize further mitochondrial genomes from slowly-evolving bryozoans to obtain a more credible lophophorate phylogeny. Results The complete mitochondrial genome (15,433 bp of Bugula neritina (Bryozoa, Gymnolaemata, Cheilostomata, one of the most widely distributed cheliostome bryozoans, is sequenced. This second bryozoan mitochondrial genome contains the set of 37 components generally observed in other metazoans, differing from that of F. hispida (Bryozoa, Gymnolaemata, Ctenostomata, which has only 36 components with loss of tRNAser(ucn genes. The B. neritina mitochondrial genome possesses 27 multiple noncoding regions. The gene order is more similar to those of the two remaining lophophorate phyla (Brachiopoda and Phoronida and a chiton Katharina tunicate than to that of F. hispida. Phylogenetic analyses based on the nucleotide sequences or amino acid residues of 12 protein-coding genes showed consistently that, within the Lophotrochozoa, the monophyly of the bryozoan class Gymnolaemata (B. neritina and F. hispida was strongly supported and the bryozoan clade was grouped with brachiopods. Echiura appeared as a subtaxon of Annelida, and Entoprocta as a sister taxon of Phoronida. The clade of Bryozoa + Brachiopoda was clustered with either the clade of Annelida-Echiura or that of Phoronida + Entoprocta. Conclusion This study presents the complete mitochondrial genome of a

  14. Main consistent patterns of Stromatoporoid Development in the Late Ordovician and Silurian in the North Urals Palaeobasin (United States)

    Antropova, E.


    In the history of the Earth there have been no basins with similar characteristics. The North Urals palaeobasin had its own unique features. The dominant benthic organisms of basin ecosystem during the Ordovician and Silurian were stromatoporoids, corals, and brachiopods. This fauna is vitally important for the aims of stratigraphy so long as conodonts are extremely rare in sections of the Northern Urals area. The most complete ordering of stromatoporoid complexes has been established and made it possible to estimate rates and measures of extinction at a level of the province. It was also found out that stromatoporoids were organisms responsive to subtle changes of environment and that they accommodated differently to those changing conditions. The evolution of stromatoporoids was accompanied by phylogenetic reorganization and formation of endemic communities in the Late Ordovician and Early Silurian. In the Late Silurian taxonomical diversity of stromatoporoids was mainly controlled by migration processes and cosmopolites with wide palaeogeographic links prevailed in the palaeobasin. Therefore palaeobasin at that time was open to stromatoporoid fauna migration which is confirmed by the occurrence of genera and species that disperse in coeval deposits of many areas, for example, Baltic States, Sweden, Ukraine (Podolia), Western Siberia, Arctic islands of Russia, Mongolia, Canada (islands). The evolution of stromatoporoid communities in the Ordovician-Silurian was intermitted by biotic crises. The analysis of stromatoporoid development helps to define crucial points of ecosystem's reorganizations coinciding with critical geological and biotic events in the history of the North Urals palaeobasin existence, as well as global events during the Ordovician and Silurian (Hirnantian Event, Ireviken Event, Lau Event). The analysis of crises indicates local dependence of stromatoporoid biodiversity on depositional environments. Large local biocenos reorganizations and biotic

  15. Stratigraphy and paleontology of Lower Permian rocks north of Cananea, northern Sonora, Mexico (United States)

    Blodgett, R.B.; Moore, T.E.; Gray, F.


    Lower Permian carbonate and overlying red bed clastic rocks are present in a 2 km2 stratigraphic window in the vicinity of Rancho La Cueva, Santa Cruz sheet (scale 1:50,000), northern Sonora, Mexico. This exposure lies unconformably beneath predominantly intermediate Upper Cretaceous volcanics yielding 40Ar/39Ar ages of 73.4?? 0.18 and 71.1 ?? 0.35 Ma. The lower part of the Permian succession consists of light- to medium-gray colored limestones of the Colina Limestone, with a minimum thickness of 235 m. Sedimentary features suggest shallow water, slightly restricted depositional environments. Although lacking observable fossils for the most part, two intervals of richly fossiliferous, silicified shell beds are present near the base and top of the Colina Limestone. The lower fauna consist mostly of gastropods and bivalves. The presence of a new microdomatid gastropod species. Glyptospira sonorensis n. sp., close to Glytospira arelela Plas, suggests a late Wolfcampian age for this horizon. The upper fauna are predominantly molluscan dominated (gastropods and bivalves), but some brachiopods (productids and the rhynchonellid genus Pontisia) are also present. Gastropod genera include Bellerophon, Warthia, Euomphalus (represented by the species, Euomphalus kaibabensis Chronic), Baylea, Worthenia, Naticopsis, Goniasma, Kinishbia, Cibecuia, and Glyptospira. The gastropods suggest a Leonardian (late Early Permian) age for this horizon, and many of the species have previously been recorded from the Supai Group and Kaibab Formation of northern and central Arizona. The Colina Limestone is conformably overlain by 11.2 m of light-gray lime mudstone and dolostone, assigned here to the Epitaph Dolomite, which in turn is succeeded by 58.8 m of red-colored sandstone and gray lime mudstone, assigned here to the Scherrer Formation. This Lower Permian succession is significant because it further strengthens the stratigraphic ties of southeastern Arizona rocks with those of northern

  16. 2010 Dry and 2009 - 2010 Wet Season Branchiopod Survey Report, Site 300

    Energy Technology Data Exchange (ETDEWEB)

    Dexter, W


    Lawrence Livermore National Laboratory (LLNL) requested that Condor Country Consulting, Inc. (CCCI) perform wet season surveys and manage the dry season sampling for listed branchiopods in two ponded locations within the Site 300 Experimental Test Site. Site 300 is located in Alameda and San Joaquin Counties, located between the Cities of Livermore and Tracy. The two pool locations have been identified for possible amphibian enhancement activities in support of the Compensation Plan for impacts tied to the Building 850 soil clean-up project. The Building 850 project design resulted in formal consultation with the U.S. Fish and Wildlife Service (USFWS) as an amendment (File 81420-2009-F-0235) to the site-wide Biological Opinion (BO) (File 1-1-02-F-0062) in the spring of 2009 and requires mitigation for the California tiger salamander (AMCA, Ambystoma californiense) and California red-legged frog (CRLF, Rana draytonii) habitat loss. Both pools contain breeding AMCA, but do not produce metamorphs due to limited hydroperiod. The pool to the southeast (Pool BC-FS-2) is the preferred site for amphibian enhancement activities, and the wetland to northwest (Pool OA-FS-1) is the alternate location for enhancement. However, prior to enhancement, LLNL has been directed by USFWS (BO Conservation Measure 17 iii) to 'conduct USFWS protocol-level branchiopod surveys to determine whether listed brachiopod species are present within the compensation area.' CCCI conducted surveys for listed branchiopods in the 2009-2010 wet season to determine the presence of federally-listed branchiopods at the two pools (previous surveys with negative findings were performed by CCCI in 2001-2002 and 2002-2003 onsite). Surveys were conducted to partially satisfy the survey requirements of the USFWS 'Interim Survey Guidelines to Permittees for Recovery Permits under Section 10(a)(1)(A) of the Endangered Species Act for the Listed Vernal Pool Branchiopods' ('Guidelines, USFWS

  17. Canada's National Building Stone: Tyndall Stone from Manitoba (United States)

    Pratt, Brian R.; Young, Graham A.; Dobrzanski, Edward P.


    burrows belonging to Thalassinoides, which were interconnected galleries likely made by arthropods after the sediment became somewhat consolidated. Slabs bearing fossils are typically avoided in construction for esthetic reasons and because some impart weaknesses. Such slabs have, however, become increasingly popular for decorative surfaces not exposed to the elements, since the macrofossil content is so striking. Most common are receptaculitids, followed by solitary rugose corals, stromatoporoid sponges, colonial rugose and tabulate corals, brachiopods, bryozoans, gastropods, cephalopods, and trilobites; echinoderm ossicles are common in the muddy matrix. Fossil relative abundances vary stratigraphically, suggesting that subtle environmental changes took place over time.

  18. Body Size Preference of Marine Animals in Relation to Extinction Selectivity (United States)

    Sriram, A.; Idgunji, S.; Heim, N. A.; Payne, J.


    Our project encompasses an extremely specific aspect in relation to the five mass extinctions in geologic history. We asked ourselves whether larger or smaller body sizes would be better suited for surviving a mass extinction. To conduct research for our project, we used the body sizes of 17,172 marine animal genera as our primary data. These animals include echinoderms, arthropods, chordates, mollusks, and brachiopods. These creatures are perfect model organisms in terms of finding data on them because they have an excellent fossil record, and are well documented. We focused on the mean body size of these animals before and after each of the five mass extinctions (end-Ordovician, Late Devonian, end-Permian, end-Triassic, and end-Cretaceous). Our hypothesis was that the average biovolume of animals increased after each of the extinctions, with the mean size being greater after than it was before. Our size data is from the Ellis & Messina Catalogue of Ostracoda and the Treatise on Invertebrate Paleontology. We obtained stratigraphic range data The Treatise and Sepkoski (2002). In our analyses, we compared the mean size of the different animal genera before and after each extinction event. We further partitioned size change across mass extinction boundaries into three categories: the surviving genera, the extinct genera, and the newly originating genera that came about after the extinction. According to our analyses, the mean sizes did not change significantly from the genera living during the stages before the extinctions and after the extinctions. From our results, we can assume that there were not enough major increases in the overall volume of the organisms to warrant a definite conclusion that extinctions lead to larger body sizes. Further support for our findings came from the T-tests in our R code. Only the Cretaceous period showed true evidence for size changing because of the extinction; in this case, the mean size decreased. T-tests for the Cretaceous

  19. Ladinian skonca beds of the Idrija Ore Deposit (W Slovenia

    Directory of Open Access Journals (Sweden)

    Jože Čar


    part of sequence C has a carbonate structure (subsequence C1 and the top part has a siliceous structure (subsequence C2, the rocks cannot be distinguished from one another by their appearance. All are strongly bituminous and shaly. Sequence C is comprised of grey to black bituminous dolomite and quartz sandstone rich in pyrite and marcasite, dolomite or quartz siltstone, and shaly claystone with coal inclusions. The sequence of C rocks ends with clayey-lythic quartz sandstone containing remains of the brachiopod Discina. These are followed by various pyroclastic rocks of sequence D. The Skonca beds and the rich bedded cinnabar ores have various sedimentary structures which accurately define the events and environments of their origin. Except for the erosion channels, which are filled with dolomite gravel, there are no other presedimentary structures in the Skonca beds (subsequence C1. However, the beds are rich in synsedimentary physical forms, particularly various stratifications, laminations and various forms of graded bedding. Among the postsedimentary structures, mention should be made of the abundant sinking and slump structures. In some places, bioturbation and the replacement of organic particles with pyrite can be observed. The Skonca beds were deposited in a spatially complex, marsh-lacustrine-lagoonal environment. The rocks of sequence A and partly also of sequence B were formed in an alkaline, oxygen-rich environment, and the lithological units of subsequence C1 in a reductive environment. The rapid transitions among rocks indicate strictly limited and slow lateral inflows of fresh water into the marshy environment overgrown with low vegetation. The rocks in subsequence C2 prove that the freshwater marsh was flooded with sea water. Deposited in the shallow, closed lagoon overgrown with low vegetation were various bituminous shales, mudstones and sandstones filled with numerous radiolarians and needles of siliceous sponges and coal intercalations. The

  20. Are we there yet? An NSF-CAREER sponsored field program as a vehicle for engaging high school students in geology (United States)

    Frank, T. D.


    little outdoor experience, to "get their hands dirty" was a major hurdle in some years. Finding the first fossil generally broke the ice, but this hurdle was never fully breached and remains a concern. At the end of the program, all participants indicated an increased appreciation for Nebraska's rich geologic heritage and for the natural sciences in general. Students went home with a collections of rocks ranging from chalk to tuff and fossils ranging from brachiopods to turtle shells. They were eager to relate Nebraska's geologic history to their parents and friends. At least five students intend to pursue a degree in geology because of their History on the Rocks experience. In 2011, the program was incorporated into the Big Red Summer Academic Camps program, a joint 4H-UNL effort. This integration allows History on the Rocks to continue after the CAREER grant expires.

  1. Brackish to hypersaline lake dolostones of the Mississippian (United States)

    Bennett, Carys; Kearsey, Timothy; Davies, Sarah; Millward, David; Marshall, John


    Flat-lying ferroan dolomite beds are common in the Mississippian and occur along the southern margin of Laurassia, from Kentucky USA to Poland. These rocks are important as they record shallow marine to coastal plain environments that may have acted as a pathway or refugia for animals that were radiating into freshwaters such as tetrapods, fish, molluscs and arthropods. This study is a contribution to the TW:eed Project (Tetrapod World: early evolution and diversification), that examines the rebuilding of Carboniferous ecosystems following a mass extinction at the end of the Devonian. The project focuses on the Tournaisian Ballagan Formation of Scotland, which contains rare fish and tetrapod fossils. The formation is characterised by an overbank facies association of siltstone, sandstone and palaeosols, interbedded with dolostone and evaporite units, and cut by fluvial sandstone facies associations of fining-upwards conglomerate lags, cross-bedded sandstone and rippled siltstone. Two sites are used as a case study to describe the sedimentological, faunal and ichnofaunal diversity of these dolostones. More than 270 dolostone beds are recorded in each of the 500 metre depth Norham Core (near Berwick-upon-Tweed) and from a 520 metre thick field section at Burnmouth. The beds are laterally extensive, over ˜1 km, although individual units do not appear to correlate between the core and the field site. In the Norham Core dolostones comprise up to 14% of the succession. 17% of the beds contain marginal marine fossils: Spirorbis, rare orthocones, brachiopods and putative marine sharks. More common fauna include ostracods, bivalves, plants, eurypterids, gastropods and sarcopterygian fish, which are interpreted as brackish to freshwater tolerant. Bioturbation is fairly common with Serpula colonies within dolostone beds and Chondrites burrowing down from the base of dolostone beds. Some rare units in the field section have a bulbous bed surface and preserve tree root traces

  2. Multiple palaeokarst horizons in the Lower Palaeozoic of Baltoscandia challenging the dogma of a deep epicontinental sea (United States)

    Lehnert, O.; Calner, M.; Ahlberg, P.; Harper, D. A.


    Several prominent palaeokarst surfaces have recently been detected in the Cambro-Ordovician sedimentary succession of Sweden. The oldest palaeokarst was found in autumn 2011 in Västergötland. An irregular palaeokarst cave with a breccia fill yielding large, angular Orsten clasts in a dark mud- to wackestone matrix is exposed beneath a karstic surface in the Cambrian Alum Shale Formation at Kakeled Quarry, Kinnekulle. The karstic surface occurs near the top of the Kakeled Limestone Bed that ranges from the upper Agnostus pisiformis into the Ctenopyge tumida Zone. The base of the cave is more than 1.4 m below this unconformity. A mass occurrence of Orusia lenticularis, a shallow-water brachiopod originally settling on hard substrates, in the karst pockets reflects deposition of the conglomeratic cover in extremely shallow marine environments. We interpret the widespread Orusia occurrences together with a brecciated or conglomeratic interval above an irregular surface in various Swedish locations as evidence for transgression after a major regression, regionally exposing the sea-floors of the Alum Shale Basin. A slightly younger karst surface is exposed in Tomten Quarry at Torbjörntorp, Västergötland. This resembles "Schrattenkalk" in the quarry wall but rock slabs cut vertical and parallel to bedding display a karren system, which reconstructed in 3D resembles "Napfkarren" or cockling features. Trilobites of the Ctenopyge bisulcata and C. linnarssoni zones have been recovered from the orsten bed just below the base of a 1-2 cm thick and irregular glauconitic packstone layer of the Bjørkåsholmen Formation (upper P. deltifer conodont Zone). The huge stratigraphic gap comprises the six uppermost trilobite zones of the Furongian plus most of the Tremadocian. Darriwilian conodonts with reworked older material within a limestone bed slightly above the glauconitic packstone point to yet another substantial gap in the succession. In the new Tingskullen core from

  3. Stratigraphy of the Silurian outcrop belt on the east side of the Cincinnati Arch in Kentucky, with revisions in the nomenclature (United States)

    McDowell, Robert C.


    Silurian rocks form a narrow arcuate outcrop belt about 100 mi long on the east side of the Cincinnati Arch in Kentucky. They range from as much as 300 ft thick in the north to a pinchout edge in the south. The nomenclature of this sequence is revised to reflect mappability and lithologic uniformity on the basis of detailed mapping at a scale of 1:24,000 by the U.S. Geological Survey in cooperation with the Kentucky Geological Survey. The Silurian rocks are divided into two parts: the Crab Orchard Group, raised in rank from Crab Orchard Formation and redefined, in the lower part of the Silurian section, and Bisher Dolomite in the upper part of the section. The Crab Orchard Group is subdivided into the Drowning Creek Formation (new name) at the base of the Silurian, overlain by the Alger Shale (adopted herein) south of Fleming County and by the Estill Shale (elevated to formational rank) north of Bath County. The Brassfield Member (reduced in rank from Brassfield Dolomite or Formation) and the Plum Creek Shale and Oldham Members of the former Crab Orchard Formation are included as members of the Drowning Creek; the Lulbegrud Shale, Waco, and Estill Shale Members of the former Crab Orchard Formation are now included in the Alger. The Drowning Creek Formation, 20 to 50 ft thick, is composed mainly of gray fine to coarse-grained dolomite with shale interbeds. The dolomite beds average several inches thick, with bedding surfaces that are locally smooth but generally irregular and are fossiliferous in many places; fossils include brachiopods, crinoid columnals, horn corals, colonial corals, trilobites, pelecypods, and bryozoans. The shale interbeds average several inches thick, except for its Plum Creek Shale Member which is entirely shale and as much as 12 ft thick, and are most abundant in the upper half of the formation. The members of the Drowning Creek intergrade and are indistinguishable in the northern part of the area. The Alger Shale, as much as 170 feet thick

  4. Geology of the Cupsuptic quadrangle, Maine (United States)

    Harwood, David S.


    The Cupsuptic quadrangle, in west-central Maine, lies in a relatively narrow belt of pre-Silurian rocks extending from the Connecticut River valley across northern New Hampshire to north-central Maine. The Albee Formation, composed of green, purple, and black phyllite with interbedded-quartzite, is exposed in the core of a regional anticlinorium overlain to the southeast by greenstone of the Oquossoc Formation which in turn is overlain by black slate of the Kamankeag Formation. In the northern part of the quadrangle the Albee Formation is overlain by black slate, feldspathic graywacke, and minor greenstone of the Dixville Formation. The Kamankeag Formation is dated as 1-ate Middle Ordovician by graptolites (zone 12) found near the base of the unit. The Dixville Formation is correlated with the Kamankeag Formation and Oquossoc Formation and is considered to be Middle Ordovician. The Albee Formation is considered to be Middle to Lower Ordovician from correlations with similar rocks in northeastern and southwestern Vermont. The Oquossoc and Kamankeag Formations are correlated with the Amonoosuc and Partridge Formations of northern New Hampshire. The pre-Silurian rocks are unconformably overlain by unnamed rocks of Silurian age in the southeast, west-central, and northwest ninths of the quadrangle. The basal Silurian units are boulder to cobble polymict conglomerate and quartz-pebble conglomerate of late Lower Silurian (Upper Llandovery) age. The overlying rocks are either well-bedded slate and quartzite, silty limestone, or arenaceous limestone. Thearenaceous limestone contains Upper Silurian (Lower Ludlow) brachiopods. The stratified rocks have been intruded by three stocks of biotite-muscovite quartz monzonite, a large body of metadiorite and associated serpentinite, smaller bodies of gabbro, granodiorite, and intrusive felsite, as well as numerous diabase and quartz monzonite dikes. The metadiorite and serpentinite, and possibly the gabbro and granodiorite are Late

  5. Lower Cretaceous Puez key-section in the Dolomites - towards the mid-Cretaceous super-greenhouse (United States)

    Lukeneder, A.; Halásová, E.; Rehákova, D.; Józsa, Š.; Soták, J.; Kroh, A.; Jovane, L.; Florindo, F.; Sprovieri, M.; Giorgioni, M.; Lukeneder, S.


    dtirol' in Bozen, Southern Tyrol. Producing major results with a broad impact requires using tools such as facies analysis supported by lithological, sedimentological and chemical characteristics, isotope and magnetic properties as well as fossil record (ammonites, belemnites, brachiopods, echinoids, planktonic foraminiferas, radiolarians, nannofossils, calcareous dinoflagellates, calpionellids). Foraminiferal study provides the zonal subdivision of the Puez section from Valanginian - Hauterivian gorbachikellids and praehedbergelids (Hedbergella semielongata Zone), Barremo-Aptian praehedbergelids (Blesusciana kuznetzove Zone), Aptian hedbergellids of occulta - aptiana - praetrocoidea group, Early Late Aptian pseudo-planispiral foraminifera (Praehedbergella luterbacheri and Globigerinelloides ferreolensis Zones), important marker species of Hedbergella trocoidea and Paraticinella bejaaouaensis for the Late Aptian zone, Early Albian microperforate hedbergellids (Hedbergella planispira Zone), Mid Albian ticinellids (Ticinella primula Zone), advanced ticinellids like Ticinella roberti etc. (Biticinella breggiensis Zone), Latest Albian rotalliporids (Rotalipora appeninica Zone) up to Early Cenonanian appearance of Thalmanninella (Rotalipora) globotruncanoides. Results of this integrated study will be used for both, the precise biostratigraphy of the sequence studied as well as for the paleoenvironmental reconstruction. Lukeneder A. 2012. New biostratigraphic data on an Upper Hauterivian-Upper Barremian ammonite assemblage from the Dolomites (Southern Alps, Italy). Cretaceous Research. doi:10.1016/j.cretres.2011.11.002 Lukeneder A. 2011. The Biancone and Rosso Ammonitico facies of the northern Trento Plateau (Dolomites, Southern Alps; Italy). Annalen des Naturhistorischen Museum Wien, Serie A, 112, 9-33. Lukeneder A. 2010. Lithostratigraphic definition and stratotype for the Puez Formation: formalisation of the Lower Cretaceous in the Dolomites (S. Tyrol, Italy). Austrian Journals

  6. Restoration of marine ecosystems following the end-Permian mass extinction: pattern and dynamics (United States)

    Chen, Z.


    Life came closest to complete annihilation during the end-Permian mass extinction (EPME). Pattern and cause of this great dying have long been disputed. Similarly, there is also some debate on the recovery rate and pattern of marine organisms in the aftermath of the EPME. Some clades recovered rapidly, within the first 1-3 Myr of the Triassic. For instance, foraminiferal recovery began 1 Myr into the Triassic and was not much affected by Early Triassic crises. Further, some earliest Triassic body and trace fossil assemblages are also more diverse than predicted. Others, ie. Brachiopods, corals etc., however, did not rebound until the Middle Triassic. In addition, although ammonoids recovered fast, reaching a higher diversity by the Smithian than in the Late Permian, much of this Early Triassic radiation was within a single group, the Ceratitina, and their morphological disparity did not expand until the end-Spathian. Here, I like to broaden the modern ecologic network model to explore the complete trophic structure of fossilized ecosystems during the Permian-Triassic transition as a means of assessing the recovery. During the Late Permian and Early Triassic, primary producers, forming the lowest trophic level, were microbes. The middle part of the food web comprises primary and meso-consumer trophic levels, the former dominated by microorganisms such as foraminifers, the latter by opportunistic communities (i.e. disaster taxa), benthic shelly communities, and reef-builders. They were often consumed by invertebrate and vertebrate predators, the top trophic level. Fossil record from South China shows that the post-extinction ecosystems were degraded to a low level and typified by primary producers or opportunistic consumers, which are represented by widespread microbialites or high-abundance, low-diversity communities. Except for some opportunists, primary consumers, namely foraminifers, rebounded in Smithian. Trace-makers recovered in Spathian, which also saw

  7. Direct Experiments on the Ocean Disposal of Fossil Fuel CO2

    Energy Technology Data Exchange (ETDEWEB)

    Barry, James, P.


    effects of high CO2 waters on marine animals (Barry et al. 2008). This system is capable of controlling oxygen, pH, and temperature of seawater for use in studies of the physiological responses of animals under acidified conditions. We have investigated the tolerance of deep- and shallow-living crabs to high CO2 levels (Pane and Barry 2007; Pane et al. 2008), and are now working on brachiopods (Barry et al. in prep.) and a comparison of deep and shallow living sea urchins. This research program, supported in part by DoE has contributed to a number of other publications authored or co-authored by Barry (Caldeira et al. 2005; Brewer and Barry 2008; Barry et al. 2006, 2010a,b,c; National Research Council, in press; Hoffman et al. in press) as well as over 40 invited talks since 2004, including Congressional briefings and testimony at U.S. Senate Hearings on Ocean Acidification. Through the grant period, the research emphasis shifted from studies of the effects of direct deep-sea carbon dioxide sequestration on deep-sea animals, to a broader conceptual framework of the effects of ocean acidification (whether purposeful or passive) on the physiology and survival of deep and shallow living marine animals. We feel that this has been a very productive program and are grateful to DoE for its support.

  8. The Cannery Formation--Devonian to Early Permian arc-marginal deposits within the Alexander Terrane, Southeastern Alaska (United States)

    Karl, Susan M.; Layer, Paul W.; Harris, Anita G.; Haeussler, Peter J.; Murchey, Benita L.


    The Cannery Formation consists of green, red, and gray ribbon chert, siliceous siltstone, graywacke-chert turbidites, and volcaniclastic sandstone. Because it contains early Permian fossils at and near its type area in Cannery Cove, on Admiralty Island in southeastern Alaska, the formation was originally defined as a Permian stratigraphic unit. Similar rocks exposed in Windfall Harbor on Admiralty Island contain early Permian bryozoans and brachiopods, as well as Mississippian through Permian radiolarians. Black and green bedded chert with subordinate lenses of limestone, basalt, and graywacke near Kake on Kupreanof Island was initially correlated with the Cannery Formation on the basis of similar lithology but was later determined to contain Late Devonian conodonts. Permian conglomerate in Keku Strait contains chert cobbles inferred to be derived from the Cannery Formation that yielded Devonian and Mississippian radiolarians. On the basis of fossils recovered from a limestone lens near Kake and chert cobbles in the Keku Strait area, the age of the Cannery Formation was revised to Devonian and Mississippian, but this revision excludes rocks in the type locality, in addition to excluding bedded chert on Kupreanof Island east of Kake that contains radiolarians of Late Pennsylvanian and early Permian age. The black chert near Kake that yielded Late Devonian conodonts is nearly contemporaneous with black chert interbedded with limestone that also contains Late Devonian conodonts in the Saginaw Bay Formation on Kuiu Island. The chert cobbles in the conglomerate in Keku Strait may be derived from either the Cannery Formation or the Saginaw Bay Formation and need not restrict the age of the Cannery Formation, regardless of their source. The minimum age of the Cannery Formation on both Admiralty Island and Kupreanof Island is constrained by the stratigraphically overlying fossiliferous Pybus Formation, of late early and early late Permian age. Because bedded radiolarian

  9. The Late Ordovician Extinction: How it became the best understood of the five major extinctions. (United States)

    Sheehan, P.


    epicontinental seas were drained in many places. An extensive record of changes of all the major faunal groups has been established and work continues. Compilations by Sepkoski and Benton established the Ordovician extinction as one of the five major Phanerozoic extinctions, ranking second only to the end Permian extinction in terms of taxonomic loss. However, as the ecologic changes caused by the extinction became better understood it was realized that of the five extinction events the Ordovician extinction caused the least ecologic perturbation. Given the interest and extensive study extinction events have generated in the last 20 years it is surprising the oldest of the five extinctions has the most well understood cause and the best global record of the faunal changes. In fact only one other extinction event (K/T event) has a widely accepted cause, darkness associated with an impact event. The general faunal changes allow at least a preliminary comparison of two events with differing causes. The most important factor promoting survival in both events is wide geographic distribution. Other ecologic factors differ considerably between the events. Extinction was very high in epicontinental seas during the Ordovician but not in the Cretaceous. Cretaceous organisms that could survive several months without food (such as animals with low metabolic rates, or larval stages that included dormancy) preferentially survived, while this was not a factor in the Ordovician when low metabolic rates of animals like brachiopods and echinoderms provided little advantage. Animals capable of feeding on detritus during the loss of sunlight preferentially survived the Cretaceous extinction, but this was not a buffer to extinction in the Ordovician.

  10. Analysis of a intra-Carixian clay horizon into carbonate platform of the Ouarsenis (Algeria): composition, dynamic and paleo-climatic implication

    Energy Technology Data Exchange (ETDEWEB)

    Benhamou, M.; Salhi, A. [Oran Univ., Faculte des Sciences de la Terre et de l' Amenagement du Territoire, Dpt. de Geologie (Algeria)


    During the Late Sinemurian a carbonate platform has developed on the Ouarsenis area (external Tell o f the Algerian Alpine belt) with setting deposits of the Kef Sidi Amar Carbonate Formation. A first maximum flooding materialized by a brachiopods (Zeilleriids) layer, is occurring during the Late Carixian. The Late Carixian deepening has been followed by a sea-level fall documented by several meters incisions filled by transgressive breccia and conglomerates. After this episode, this material was sealed by a pedogenic bed (0,05 to 0,20 m) which corresponds to a yellow clay deposit containing well rounded particles interpreted as pedo-genetic globules. These corpuscles are composed of reddish and hardened clay, corroded quartz grains, rhombic and zoned dolomite crystals and ankerite, monocrystalline and xeno-morphous detrital quartz grains (1-2 mm). The observed characteristics allow to recognize a typical calcrete. They are the result of pedo-genetic diagenesis developed inside the phreatic water-table near the surface: this is an alteration profile. The mineralogic fraction has been analyzed by X-Ray which show results of association clay mineral as a predominance of illite (85%) and mixed-layer illite-montmorillonite (I-M, 10%) associated with a low ration of chlorite (5%) and kaolinite trace (1%). This mineralogic clay association indicates a shallow water (hydro-morphic zone). Among these clay minerals, the illite reveals the precious indications in a source area. In this case, it comes from the decomposition of the schist paleo-relief located in the internal domain. This rock was transformed by acid leaching (action of the sour humus) into kaolinite with the presence of the quartzification. The origin of the mixed-layer clay I-M (10%) is the result of the active pedogenesis. The simultaneous presence of the illite, chlorite, kaolinite and the mixed-layer clay I-M seems to be result from the erosion exercised on the alteration product or arenitisation of the

  11. Sedimentology, conodonts and ostracods of the Devonian - Carboniferous strata of the Anseremme railway bridge section, Dinant Basin, Belgium (United States)

    Casier, J.-G.; Mamet, B.; Preat, A.; Sandberg, C.A.


    Seven major carbonate microfacies are defined in the Devonian - Carboniferous (D/C) strata (50 m) of the Anseremme railway bridge section, south of Dinant. They permit recognition of several levels encompassing the Etroeungt and Hastie??re formations. "Bathymetric" sequences range from open marine, below the storm wave base, to semi-restricted lagoon. This sequence records a shallowing-upward trend of the relative sea level, from environments below the storm wave base to strongly eroded supraticial pre-evaporitic environments. Faunal components (echinoderms, brachiopods...) indicate open-marine domain for the first six microfacies located within the dysphoticeuphotic zone in relatively shallow waters. The textures of the rocks (mudstones to rudstones) associated with lamination characteristics indicate the position of the storm (SWB) and the fair-weather (FWWB) wave bases. Microfacies seven suggests a semi-restricted platform with salinity fluctuations from hypersaline brines to brackish waters. Thus, the boundary of the Etroeungt/Hastie??re formations is marked by an abrupt drop in sea level. Carbonate micro-conglomerates recording an important erosive phase and a sedimentary hiatus. The environment is again open marine in the upper part of the Hastie??re Formation. Our conclusion is that the Anseremme section is not a reliable continuous succession for the study of the D/C boundary. This confirms the VAN STEENWINKEL (1988, 1993 hypothesis based on other arguments. Conodont faunas demonstrate that the Devonian sequence spans the five youngest conodont zones, but that two of these zones are not represented. The Epinette Formation is dated as the youngest part of the Middle expansa Zone. Thus, the boundary with the Late praesulcata Zone probably coincides with the sharp sedimentological change at the base of the Etroeungt Formation, which is interpreted to belong entirely to this zone. The disconformably overlying basal bed 159 of the Hastie??re Formation is dated

  12. Grainstones and cementstone mounds: The Trogkofel summit section (Lower Permian, Carnic Alps, Austria). (United States)

    Schaffhauser, M.; Sanders, D.; Krainer, K.


    are overgrown by abundant, thick fringes and botryoids of fibrous cement that is interpreted as calcitized aragonite cement. In addition, brachiopods, crustose red algae, and a few solitary and colonial rugose corals are typical. By volume, the former aragonite cement comprises the majority of the mounds. Intrinsic pores within the cementstone fabrics typically are filled by micropeloidal grainstone and/or by lime mudstone. The Trogkofel Limestone is locally dolomitized. Replacement dolomites show a wide range of crystal shapes and textures, but overall comprise (a) finely-crystalline, limpid dolostone of xenotopic or hypidiotopic fabrics that broadly mimick the texture of replaced sediment and cements, (b) coarse-crystalline fabrics of hypidiotopic to idiotopic, limpid or optically zoned dolomite, and (c) replacement saddle dolomite. The Trogkofel Limestone is riddled by karstic dykes and caverns that are mainly filled by, both or either of, geopetally-laminated red lime mudstone, terrigenous red sandstones, or thick fringes of fibrous cement. In the karstic cavity fills, packages of convolute geopetal lamination and brecciated internal sediments (internal seismites) overlain by infills with non-convolute lamination, fracture of fibrous cements, and dykes filled by multi-phase fracture breccias record tectonism during or after deposition of the Trogkofel Limestone. The Trogkofel Limestone is capped by a truncation surface which, in turn, is overlain by an interval of extremely poorly sorted, thick-bedded breccias with a former matrix of lime mudstone ("Trogkofel Breccia"). Both the components and the matrix of the Trogkofel Breccia are dolomitized. We interpret the facies and facies architecture of the eastern cliff section of Trogkofel as succession from the seaward side of a "grainstone-dominated" platform margin with cementstone mounds. The lack of clear-cut vertical trends in prevalent facies suggests that the platform margin developed mainly by aggradation. The

  13. Diagenetic evolution and stable isotopes of Lower Permian platform marginal carbonates (Trogkofel Limestone, Carnic Alps, Austria) (United States)

    Schaffhauser, Maria; Krainer, Karl; Sanders, Diethard Gerald; Spötl, Christoph


    and filling of fractures. Closely below the erosional surface at the top of the Trogkofel Limestone, the dolomite is characterized by vuggy porosity. The Tarvis Breccia, which represents coarse alluvial fan deposits, in turn, is thick-bedded, poorly sorted, typically clast-supported, and consists of angular lithoclasts embedded in a matrix of former lime mudstone. Both, matrix as well as lithoclasts, are dolomitized. Various types of cement (isopachous, botryoidal, microbialite, calcite spar), karstic cavity fills (isopachous cements, internal sediments), and replacement dolomites of the Trogkofel section, as well as of the Tarvis Breccia were analysed for their stable isotopic composition. δ18O and δ13C data produced so far allow to differentiate between replacement dolomites and saddle dolomite of the Trogkofel Limestone and the Tarvis Breccia. Saddle dolomite shows the most depleted oxygen isotope values, suggesting formation during relatively high temperatures. Carbon isotope values are invariably positive in all dolomite types indicating lacking influence of organic diagenesis on the alkalinity of the deep-burial pore water. Matrix dolomite from the Tarvis Breccia shows slightly positive δ18O values. Calcite cements show a wide range in δ18O values (ca. -1 to -7 permil VPDB), which overlaps the composition of unaltered brachiopod shells (ca. -3 permil VPDB). Oxygen isotope values of calcite cements reveal a trend towards depleted δ18O values. This trend is reflecting most likely increasing temperature.

  14. Petrography, Geochemistry and Proposed Genesis of Ordovician Oolitic Iron Formation Members of the Lashkarak Formation, Eastern Alborz

    Directory of Open Access Journals (Sweden)

    Mansoore Maghsoudloo Mahalli


    , Iran. Logistical and financial support was provided by the Research Grant to senior author. We are grateful to SarCheshmeh Copper Complex for XRF analyses and IMPERC for XRD measurements. We gratefully acknowledge Dr. Ghobadipour for SEM analysis in National Museum of Wales, Great Britain. References Ghobadi Pour, M., Popov, L.E., Kebriaee-Zadeh, M.R. and Baars C.H., 2011. Middle Ordovician (Darriwilian Brachiopods associated with the Neseuretus bio-facies, Eastern Alborz Mountains, Iran. Memoirs of the Association of Australasian Palaeontologists, 42(3: 263-283. Maynard, J.B., 1983. Geochemistry of sedimentary ore deposits. Springer-Verlag, NewYork, 382 pp. Maynard, J.B., 1986. Geochemistry of oolitic iron ores, an electron microprobe study. Economic Geology, 81(8: 1473-1483. Mucke, A.T. and Farshad, F., 2005. Whole-rock and mineralogical composition of Phanerozoic ooidal ironstones: Comparison and differentiation of types and subtypes. Ore Geology Reviews, 26(2: 227-262. Petranek, J. and Van Houton, F.B., 1997. Phanerozoic ooidal ironstone. Czech Geological Survey, Special Papers 7: 70 pp. Young, T.P., 1989. Eustatically controlled ooidal ironstone deposition: facies relationships of the Ordovician open-shelf ironstones of Western Europe. In: T.P. Young and W.E.G. Taylor (Editors, Phanerozoic Ironstones. Geological Society of London, Special Publication, 46(1: 51–64.

  15. Siberia, the wandering northern terrane, and its changing geography through the Palaeozoic (United States)

    Cocks, L. Robin M.; Torsvik, Trond H.


    The old terrane of Siberia occupied a very substantial area in the centre of today's political Siberia and also adjacent areas of Mongolia, eastern Kazakhstan, and northwestern China. Siberia's location within the Early Neoproterozoic Rodinia Superterrane is contentious (since few if any reliable palaeomagnetic data exist between about 1.0 Ga and 540 Ma), but Siberia probably became independent during the breakup of Rodinia soon after 800 Ma and continued to be so until very near the end of the Palaeozoic, when it became an integral part of the Pangea Supercontinent. The boundaries of the cratonic core of the Siberian Terrane (including the Patom area) are briefly described, together with summaries of some of the geologically complex surrounding areas, and it is concluded that all of the Palaeozoic underlying the West Siberian Basin (including the Ob-Saisan Surgut area), Tomsk Terrane, Altai-Sayan Terranes (including Salair, Kuznetsk Alatau, Batenov, Kobdin and West Sayan), Ertix Terrane, Barguzin Terrane, Tuva-Mongol Terrane, Central Mongolia Terrane Assemblage, Gobi Altai and Mandalovoo Terranes, Okhotsk Terrane and much of the Verkhoyansk-Kolyma region all formed parts of peri-Siberia, and thus rotated with the main Siberian Craton as those areas were progressively accreted to the main Siberian Terrane at various times during the latest Neoproterozoic and Palaeozoic. The Ertix Terrane is a new term combining what has been termed the "Altay Terrane" or "NE Xinjiang" area of China, and the Baytag, Baaran and Bidz terranes of Mongolia. The Silurian Tuvaella brachiopod fauna is restricted only to today's southern parts of peri-Siberia. Thus, allowing for subsequent rotation, the fauna occurs only in the N of the Siberian Terrane, and, as well as being a helpful indicator of what marginal terranes made up peri-Siberia, is distinctive as being the only Silurian fauna known from northern higher latitudes globally. In contrast, the other terranes adjacent to peri

  16. Comparative EPMA and μXRF methods for mapping micro-scale distribution of iodine in bio-carbonates of the Callovian-Oxfordian clayey formation at Bure, Eastern part of the Paris Basin

    International Nuclear Information System (INIS)

    Lerouge, C.; Claret, F.; Wille, G.; Beny, C.; Gaucher, E.C.; Tournassat, C.; Denecke, M.A.; Falkenberg, G.; Ramboz, C.; Giffaut, E.


    Document available in extended abstract form only. A pluri-disciplinary approach was applied on carbonate shells of the Callovian-Oxfordian (160 Ma) clayey formation from the ANDRA Underground Research Laboratory at Bure. The overall aim of this natural analog investigation was to define the iodine speciation and immobilization mechanisms by carbonates in a natural carbonate-bearing clayey formation. In this study, sequential chemical extraction data have shown that ∼66 % of the low iodine content of the clay stone (1-5 ppm) is associated with the carbonate fraction and ∼33 % with organic matter. EPMA analyses have been performed on detrital and different generations of diagenetic carbonates observed in thin sections and have provided evidence of low and heterogeneously distributed iodine contents measured in bio-clasts (up to 1300 ppm), but also in less amounts in diagenetic euhedral calcite and dolomite (up to 620 ppm). EPMA analyses to measure iodine is especially challenging due to spectral interference between calcium K and iodine L emission lines and to the relatively high detection limits (∼150 ppm). We therefore applied a synchrotron micro-focused beam at energies above the I K absorption in the present study. Two entire and centimeter-sized shells, a bivalve and a brachiopod (Rhynchonella) were separated in a clay stone sample from Gallery level of the URL at Bure. They were cut along two perpendicular directions to obtain thin polished sections of the bio-clast shells. A dry polishing procedure was performed to avoid any partial dissolution of calcite and done by hand to avoid any heating and volatilization of iodine. Combined results from optical microscopy, SEM, cathodoluminescence and infra-red spectrometry performed on both bio-clasts showed that the Rhynchonella shell has a well-preserved organic structure and dominantly consists of bio-calcite, whereas the bivalve shell is entirely recrystallized into diagenetic calcite, with inclusions of